Chapter VI: Book III: Modern Development of the Physical Sciences (3)

Since then star-charting with the film has come virtually to supersede the old method. A concerted effort is being made by astronomers in various parts of the world to make a complete chart of the heavens, and before the close of our century this work will be accomplished, some fifty or sixty millions of visible stars being placed on record with a degree of accuracy hitherto unapproachable. Moreover, other millions of stars are brought to light by the negative, which are too distant or dim to be visible with any telescopic powers yet attained--a fact which wholly discredits all previous inferences as to the limits of our sidereal system. Hence, notwithstanding the wonderful instrumental advances of the nineteenth century, knowledge of the exact form and extent of our universe seems more unattainable than it seemed a century ago.

The Structure of Nebulae

Yet the new instruments, while leaving so much untold, have revealed some vastly important secrets of cosmic structure. In particular, they have set at rest the long-standing doubts as to the real structure and position of the mysterious nebulae--those lazy masses, only two or three of them visible to the unaided eye, which the telescope reveals in almost limitless abundance, scattered everywhere among the stars, but grouped in particular about the poles of the stellar stream or disk which we call the Milky Way.

Herschel's later view, which held that some at least of the nebulae are composed of a "shining fluid," in process of condensation to form stars, was generally accepted for almost half a century. But in 1844, when Lord Rosse's great six-foot reflector--the largest telescope ever yet constructed--was turned on the nebulae, it made this hypothesis seem very doubtful. Just as Galileo's first lens had resolved the Milky Way into stars, just as Herschel had resolved nebulae that resisted all instruments but his own, so Lord Rosse's even greater reflector resolved others that would not yield to Herschel's largest mirror. It seemed a fair inference that with sufficient power, perhaps some day to be attained, all nebulae would yield, hence that all are in reality what Herschel had at first thought them--vastly distant "island universes," composed of aggregations of stars, comparable to our own galactic system.

But the inference was wrong; for when the spectroscope was first applied to a nebula in 1864, by Dr. Huggins, it clearly showed the spectrum not of discrete stars, but of a great mass of glowing gases, hydrogen among others. More extended studies showed, it is true, that some nebulae give the continuous spectrum of solids or liquids, but the different types intermingle and grade into one another. Also, the closest affinity is shown between nebulae and stars. Some nebulae are found to contain stars, singly or in groups, in their actual midst; certain condensed "planetary" nebulae are scarcely to be distinguished from stars of the gaseous type; and recently the photographic film has shown the presence of nebulous matter about stars that to telescopic vision differ in no respect from the generality of their fellows in the galaxy. The familiar stars of the Pleiades cluster, for example, appear on the negative immersed in a hazy blur of light. All in all, the accumulated impressions of the photographic film reveal a prodigality of nebulous matter in the stellar system not hitherto even conjectured.

And so, of course, all question of "island universes" vanishes, and the nebulae are relegated to their true position as component parts of the one stellar system--the one universe--that is open to present human inspection. And these vast clouds of world-stuff have been found by Professor Keeler, of the Lick observatory, to be floating through space at the starlike speed of from ten to thirty-eight miles per second.

The linking of nebulae with stars, so clearly evidenced by all these modern observations, is, after all, only the scientific corroboration of what the elder Herschel's later theories affirmed. But the nebulae have other affinities not until recently suspected; for the spectra of some of them are practically identical with the spectra of certain comets. The conclusion seems warranted that comets are in point of fact minor nebulae that are drawn into our system; or, putting it otherwise, that the telescopic nebulae are simply gigantic distant comets.

Lockyer's Meteoric Hypothesis

Following up the surprising clews thus suggested, Sir Norman Lockyer, of London, has in recent years elaborated what is perhaps the most comprehensive cosmogonic guess that has ever been attempted. His theory, known as the "meteoric hypothesis," probably bears the same relation to the speculative thought of our time that the nebular hypothesis of Laplace bore to that of the eighteenth century. Outlined in a few words, it is an attempt to explain all the major phenomena of the universe as due, directly or indirectly, to the gravitational impact of such meteoric particles, or specks of cosmic dust, as comets are composed of. Nebulae are vast cometary clouds, with particles more or less widely separated, giving off gases through meteoric collisions, internal or external, and perhaps glowing also with electrical or phosphorescent light. Gravity eventually brings the nebular particles into closer aggregations, and increased collisions finally vaporize the entire mass, forming planetary nebulae and gaseous stars. Continued condensation may make the stellar mass hotter and more luminous for a time, but eventually leads to its liquefaction, and ultimate consolidation--the aforetime nebulae becoming in the end a dark or planetary star.

The exact correlation which Lockyer attempts to point out between successive stages of meteoric condensation and the various types of observed stellar bodies does not meet with unanimous acceptance. Mr. Ranyard, for example, suggests that the visible nebulae may not be nascent stars, but emanations from stars, and that the true pre-stellar nebulae are invisible until condensed to stellar proportions. But such details aside, the broad general hypothesis that all the bodies of the universe are, so to speak, of a single species--that nebulae (including comets), stars of all types, and planets, are but varying stages in the life history of a single race or type of cosmic organisms--is accepted by the dominant thought of our time as having the highest warrant of scientific probability.

All this, clearly, is but an amplification of that nebular hypothesis which, long before the spectroscope gave us warrant to accurately judge our sidereal neighbors, had boldly imagined the development of stars out of nebulae and of planets out of stars. But Lockyer's hypothesis does not stop with this. Having traced the developmental process from the nebular to the dark star, it sees no cause to abandon this dark star to its fate by assuming, as the original speculation assumed, that this is a culminating and final stage of cosmic existence. For the dark star, though its molecular activities have come to relative stability and impotence, still retains the enormous potentialities of molar motion; and clearly, where motion is, stasis is not. Sooner or later, in its ceaseless flight through space, the dark star must collide with some other stellar body, as Dr. Croll imagines of the dark bodies which his "pre-nebular theory" postulates. Such collision may be long delayed; the dark star may be drawn in comet-like circuit about thousands of other stellar masses, and be hurtled on thousands of diverse parabolic or elliptical orbits, before it chances to collide--but that matters not: "billions are the units in the arithmetic of eternity," and sooner or later, we can hardly doubt, a collision must occur. Then without question the mutual impact must shatter both colliding bodies into vapor, or vapor combined with meteoric fragments; in short, into a veritable nebula, the matrix of future worlds. Thus the dark star, which is the last term of one series of cosmic changes, becomes the first term of another series--at once a post-nebular and a pre-nebular condition; and the nebular hypothesis, thus amplified, ceases to be a mere linear scale, and is rounded out to connote an unending series of cosmic cycles, more nearly satisfying the imagination.

In this extended view, nebulae and luminous stars are but the infantile and adolescent stages of the life history of the cosmic individual; the dark star, its adult stage, or time of true virility. Or we may think of the shrunken dark star as the germ-cell, the pollen-grain, of the cosmic organism. Reduced in size, as becomes a germ-cell, to a mere fraction of the nebular body from which it sprang, it yet retains within its seemingly non-vital body all the potentialities of the original organism, and requires only to blend with a fellow-cell to bring a new generation into being. Thus may the cosmic race, whose aggregate census makes up the stellar universe, be perpetuated--individual solar systems, such as ours, being born, and growing old, and dying to live again in their descendants, while the universe as a whole maintains its unified integrity throughout all these internal mutations--passing on, it may be, by infinitesimal stages, to a culmination hopelessly beyond human comprehension.

III. THE NEW SCIENCE OF PALEONTOLOGY

WILLIAM SMITH AND FOSSIL SHELLS

Ever since Leonardo da Vinci first recognized the true character of fossils, there had been here and there a man who realized that the earth's rocky crust is one gigantic mausoleum. Here and there a dilettante had filled his cabinets with relics from this monster crypt; here and there a philosopher had pondered over them--questioning whether perchance they had once been alive, or whether they were not mere abortive souvenirs of that time when the fertile matrix of the earth was supposed to have

"teemed at a birth
Innumerous living creatures, perfect forms,
Limbed and full grown."

Some few of these philosophers--as Robert Hooke and Steno in the seventeenth century, and Moro, Leibnitz, Buffon, Whitehurst, Werner, Hutton, and others in the eighteenth--had vaguely conceived the importance of fossils as records of the earth's ancient history, but the wisest of them no more suspected the full import of the story written in the rocks than the average stroller in a modern museum suspects the meaning of the hieroglyphs on the case of a mummy.

It was not that the rudiments of this story are so very hard to decipher--though in truth they are hard enough--but rather that the men who made the attempt had all along viewed the subject through an atmosphere of preconception, which gave a distorted image. Before this image could be corrected it was necessary that a man should appear who could see without prejudice, and apply sound common-sense to what he saw. And such a man did appear towards the close of the century, in the person of William Smith, the English surveyor. He was a self-taught man, and perhaps the more independent for that, and he had the gift, besides his sharp eyes and receptive mind, of a most tenacious memory. By exercising these faculties, rare as they are homely, he led the way to a science which was destined, in its later developments, to shake the structure of established thought to its foundations.

Little enough did William Smith suspect, however, that any such dire consequences were to come of his act when he first began noticing the fossil shells that here and there are to be found in the stratified rocks and soils of the regions over which his surveyor's duties led him. Nor, indeed, was there anything of such apparent revolutionary character in the facts which he unearthed; yet in their implications these facts were the most disconcerting of any that had been revealed since the days of Copernicus and Galileo. In its bald essence, Smith's discovery was simply this: that the fossils in the rocks, instead of being scattered haphazard, are arranged in regular systems, so that any given stratum of rock is labelled by its fossil population; and that the order of succession of such groups of fossils is always the same in any vertical series of strata in which they occur. That is to say, if fossil A underlies fossil B in any given region, it never overlies it in any other series; though a kind of fossils found in one set of strata may be quite omitted in another. Moreover, a fossil once having disappeared never reappears in any later stratum.

From these novel facts Smith drew the commonsense inference that the earth had had successive populations of creatures, each of which in its turn had become extinct. He partially verified this inference by comparing the fossil shells with existing species of similar orders, and found that such as occur in older strata of the rocks had no counterparts among living species. But, on the whole, being eminently a practical man, Smith troubled himself but little about the inferences that might be drawn from his facts. He was chiefly concerned in using the key he had discovered as an aid to the construction of the first geological map of England ever attempted, and he left to others the untangling of any snarls of thought that might seem to arise from his discovery of the succession of varying forms of life on the globe.

He disseminated his views far and wide, however, in the course of his journeyings--quite disregarding the fact that peripatetics went out of fashion when the printing-press came in--and by the beginning of the nineteenth century he had begun to have a following among the geologists of England. It must not for a moment be supposed, however, that his contention regarding the succession of strata met with immediate or general acceptance. On the contrary, it was most bitterly antagonized. For a long generation after the discovery was made, the generality of men, prone as always to strain at gnats and swallow camels, preferred to believe that the fossils, instead of being deposited in successive ages, had been swept all at once into their present positions by the current of a mighty flood--and that flood, needless to say, the Noachian deluge. Just how the numberless successive strata could have been laid down in orderly sequence to the depth of several miles in one such fell cataclysm was indeed puzzling, especially after it came to be admitted that the heaviest fossils were not found always at the bottom; but to doubt that this had been done in some way was rank heresy in the early days of the nineteenth century.

CUVIER AND FOSSIL VERTEBRATES

But once discovered, William Smith's unique facts as to the succession of forms in the rocks would not down. There was one most vital point, however, regarding which the inferences that seem to follow from these facts needed verification--the question, namely, whether the disappearance of a fauna from the register in the rocks really implies the extinction of that fauna. Everything really depended upon the answer to that question, and none but an accomplished naturalist could answer it with authority. Fortunately, the most authoritative naturalist of the time, George Cuvier, took the question in hand--not, indeed, with the idea of verifying any suggestion of Smith's, but in the course of his own original studies--at the very beginning of the century, when Smith's views were attracting general attention.

Cuvier and Smith were exact contemporaries, both men having been born in 1769, that "fertile year" which gave the world also Chateaubriand, Von Humboldt, Wellington, and Napoleon. But the French naturalist was of very different antecedents from the English surveyor. He was brilliantly educated, had early gained recognition as a scientist, and while yet a young man had come to be known as the foremost comparative anatomist of his time. It was the anatomical studies that led him into the realm of fossils. Some bones dug out of the rocks by workmen in a quarry were brought to his notice, and at once his trained eye told him that they were different from anything he had seen before. Hitherto such bones, when not entirely ignored, had been for the most part ascribed to giants of former days, or even to fallen angels. Cuvier soon showed that neither giants nor angels were in question, but elephants of an unrecognized species. Continuing his studies, particularly with material gathered from gypsum beds near Paris, he had accumulated, by the beginning of the nineteenth century, bones of about twenty-five species of animals that he believed to be different from any now living on the globe.

The fame of these studies went abroad, and presently fossil bones poured in from all sides, and Cuvier's conviction that extinct forms of animals are represented among the fossils was sustained by the evidence of many strange and anomalous forms, some of them of gigantic size. In 1816 the famous Ossements Fossiles, describing these novel objects, was published, and vertebrate paleontology became a science. Among other things of great popular interest the book contained the first authoritative description of the hairy elephant, named by Cuvier the mammoth, the remains of which bad been found embedded in a mass of ice in Siberia in 1802, so wonderfully preserved that the dogs of the Tungusian fishermen actually ate its flesh. Bones of the same species had been found in Siberia several years before by the naturalist Pallas, who had also found the carcass of a rhinoceros there, frozen in a mud-bank; but no one then suspected that these were members of an extinct population--they were supposed to be merely transported relics of the flood.

Cuvier, on the other hand, asserted that these and the other creatures he described had lived and died in the region where their remains were found, and that most of them have no living representatives upon the globe. This, to be sure, was nothing more than William Smith had tried all along to establish regarding lower forms of life; but flesh and blood monsters appeal to the imagination in a way quite beyond the power of mere shells; so the announcement of Cuvier's discoveries aroused the interest of the entire world, and the Ossements Fossiles was accorded a popular reception seldom given a work of technical science--a reception in which the enthusiastic approval of progressive geologists was mingled with the bitter protests of the conservatives.

"Naturalists certainly have neither explored all the continents," said Cuvier, "nor do they as yet even know all the quadrupeds of those parts which have been explored. New species of this class are discovered from time to time; and those who have not examined with attention all the circumstances belonging to these discoveries may allege also that the unknown quadrupeds, whose fossil bones have been found in the strata of the earth, have hitherto remained concealed in some islands not yet discovered by navigators, or in some of the vast deserts which occupy the middle of Africa, Asia, the two Americas, and New Holland.

"But if we carefully attend to the kind of quadrupeds that have been recently discovered, and to the circumstances of their discovery, we shall easily perceive that there is very little chance indeed of our ever finding alive those which have only been seen in a fossil state.

"Islands of moderate size, and at a considerable distance from the large continents, have very few quadrupeds. These must have been carried to them from other countries. Cook and Bougainville found no other quadrupeds besides hogs and dogs in the South Sea Islands; and the largest quadruped of the West India Islands, when first discovered, was the agouti, a species of the cavy, an animal apparently between the rat and the rabbit.

"It is true that the great continents, as Asia, Africa, the two Americas, and New Holland, have large quadrupeds, and, generally speaking, contain species common to each; insomuch, that upon discovering countries which are isolated from the rest of the world, the animals they contain of the class of quadruped were found entirely different from those which existed in other countries. Thus, when the Spaniards first penetrated into South America, they did not find it to contain a single quadruped exactly the same with those of Europe, Asia, and Africa. The puma, the jaguar, the tapir, the capybara, the llama, or glama, and vicuna, and the whole tribe of sapajous, were to them entirely new animals, of which they had not the smallest idea....

"If there still remained any great continent to be discovered, we might perhaps expect to be made acquainted with new species of large quadrupeds, among which some might be found more or less similar to those of which we find the exuviae in the bowels of the earth. But it is merely sufficient to glance the eye over the maps of the world and observe the innumerable directions in which navigators have traversed the ocean, in order to be satisfied that there does not remain any large land to be discovered, unless it may be situated towards the Antarctic Pole, where eternal ice necessarily forbids the existence of animal life."(1)

Cuvier then points out that the ancients were well acquainted with practically all the animals on the continents of Europe, Asia, and Africa now known to scientists. He finds little grounds, therefore, for belief in the theory that at one time there were monstrous animals on the earth which it was necessary to destroy in order that the present fauna and men might flourish. After reviewing these theories and beliefs in detail, he takes up his Inquiry Respecting the Fabulous Animals of the Ancients. "It is easy," he says, "to reply to the foregoing objections, by examining the descriptions that are left us by the ancients of those unknown animals, and by inquiring into their origins. Now that the greater number of these animals have an origin, the descriptions given of them bear the most unequivocal marks; as in almost all of them we see merely the different parts of known animals united by an unbridled imagination, and in contradiction to every established law of nature."(2)

Having shown how the fabulous monsters of ancient times and of foreign nations, such as the Chinese, were simply products of the imagination, having no prototypes in nature, Cuvier takes up the consideration of the difficulty of distinguishing the fossil bones of quadrupeds.

We shall have occasion to revert to this part of Cuvier's paper in another connection. Here it suffices to pass at once to the final conclusion that the fossil bones in question are the remains of an extinct fauna, the like of which has no present-day representation on the earth. Whatever its implications, this conclusion now seemed to Cuvier to be fully established.

In England the interest thus aroused was sent to fever-heat in 1821 by the discovery of abundant beds of fossil bones in the stalagmite-covered floor of a cave at Kirkdale, Yorkshire which went to show that England, too, had once had her share of gigantic beasts. Dr. Buckland, the incumbent of the chair of geology at Oxford, and the most authoritative English geologist of his day, took these finds in hand and showed that the bones belonged to a number of species, including such alien forms as elephants, rhinoceroses, hippopotami, and hyenas. He maintained that all of these creatures had actually lived in Britain, and that the caves in which their bones were found had been the dens of hyenas.

The claim was hotly disputed, as a matter of course. As late as 1827 books were published denouncing Buckland, doctor of divinity though he was, as one who had joined in an "unhallowed cause," and reiterating the old cry that the fossils were only remains of tropical species washed thither by the deluge. That they were found in solid rocks or in caves offered no difficulty, at least not to the fertile imagination of Granville Penn, the leader of the conservatives, who clung to the old idea of Woodward and Cattcut that the deluge had dissolved the entire crust of the earth to a paste, into which the relics now called fossils had settled. The caves, said Mr. Penn, are merely the result of gases given off by the carcasses during decomposition--great air-bubbles, so to speak, in the pasty mass, becoming caverns when the waters receded and the paste hardened to rocky consistency.

But these and such-like fanciful views were doomed even in the day of their utterance. Already in 1823 other gigantic creatures, christened ichthyosaurus and plesiosaurus by Conybeare, had been found in deeper strata of British rocks; and these, as well as other monsters whose remains were unearthed in various parts of the world, bore such strange forms that even the most sceptical could scarcely hope to find their counterparts among living creatures. Cuvier's contention that all the larger vertebrates of the existing age are known to naturalists was borne out by recent explorations, and there seemed no refuge from the conclusion that the fossil records tell of populations actually extinct. But if this were admitted, then Smith's view that there have been successive rotations of population could no longer be denied. Nor could it be in doubt that the successive faunas, whose individual remains have been preserved in myriads, representing extinct species by thousands and tens of thousands, must have required vast periods of time for the production and growth of their countless generations.

As these facts came to be generally known, and as it came to be understood in addition that the very matrix of the rock in which fossils are imbedded is in many cases one gigantic fossil, composed of the remains of microscopic forms of life, common-sense, which, after all, is the final tribunal, came to the aid of belabored science. It was conceded that the only tenable interpretation of the record in the rocks is that numerous populations of creatures, distinct from one another and from present forms, have risen and passed away; and that the geologic ages in which these creatures lived were of inconceivable length. The rank and file came thus, with the aid of fossil records, to realize the import of an idea which James Hutton, and here and there another thinker, had conceived with the swift intuition of genius long before the science of paleontology came into existence. The Huttonian proposition that time is long had been abundantly established, and by about the close of the first third of the last century geologists had begun to speak of "ages" and "untold aeons of time" with a familiarity which their predecessors had reserved for days and decades.

CHARLES LYELL COMBATS CATASTROPHISM

And now a new question pressed for solution. If the earth has been inhabited by successive populations of beings now extinct, how have all these creatures been destroyed? That question, however, seemed to present no difficulties. It was answered out of hand by the application of an old idea. All down the centuries, whatever their varying phases of cosmogonic thought, there had been ever present the idea that past times were not as recent times; that in remote epochs the earth had been the scene of awful catastrophes that have no parallel in "these degenerate days." Naturally enough, this thought, embalmed in every cosmogonic speculation of whatever origin, was appealed to in explanation of the destruction of these hitherto unimagined hosts, which now, thanks to science, rose from their abysmal slumber as incontestable, but also as silent and as thought-provocative, as Sphinx or pyramid. These ancient hosts, it was said, have been exterminated at intervals of odd millions of years by the recurrence of catastrophes of which the Mosaic deluge is the latest, but perhaps not the last.

This explanation had fullest warrant of scientific authority. Cuvier had prefaced his classical work with a speculative disquisition whose very title (Discours sur les Revolutions du Globe) is ominous of catastrophism, and whose text fully sustains the augury. And Buckland, Cuvier's foremost follower across the Channel, had gone even beyond the master, naming the work in which he described the Kirkdale fossils, Reliquiae Diluvianae, or Proofs of a Universal Deluge.

Both these authorities supposed the creatures whose remains they studied to have perished suddenly in the mighty flood whose awful current, as they supposed, gouged out the modern valleys and hurled great blocks of granite broadcast over the land. And they invoked similar floods for the extermination of previous populations.

It is true these scientific citations had met with only qualified approval at the time of their utterance, because then the conservative majority of mankind did not concede that there had been a plurality of populations or revolutions; but now that the belief in past geologic ages had ceased to be a heresy, the recurring catastrophes of the great paleontologists were accepted with acclaim. For the moment science and tradition were at one, and there was a truce to controversy, except indeed in those outlying skirmish-lines of thought whither news from headquarters does not permeate till it has become ancient history at its source.

The truce, however, was not for long. Hardly had contemporary thought begun to adjust itself to the conception of past ages of incomprehensible extent, each terminated by a catastrophe of the Noachian type, when a man appeared who made the utterly bewildering assertion that the geological record, instead of proving numerous catastrophic revolutions in the earth's past history, gives no warrant to the pretensions of any universal catastrophe whatever, near or remote.

This iconoclast was Charles Lyell, the Scotchman, who was soon to be famous as the greatest geologist of his time. As a young man he had become imbued with the force of the Huttonian proposition, that present causes are one with those that produced the past changes of the globe, and he carried that idea to what he conceived to be its logical conclusion. To his mind this excluded the thought of catastrophic changes in either inorganic or organic worlds.

But to deny catastrophism was to suggest a revolution in current thought. Needless to say, such revolution could not be effected without a long contest. For a score of years the matter was argued pro and con., often with most unscientific ardor. A mere outline of the controversy would fill a volume; yet the essential facts with which Lyell at last established his proposition, in its bearings on the organic world, may be epitomized in a few words. The evidence which seems to tell of past revolutions is the apparently sudden change of fossils from one stratum to another of the rocks. But Lyell showed that this change is not always complete. Some species live on from one alleged epoch into the next. By no means all the contemporaries of the mammoth are extinct, and numerous marine forms vastly more ancient still have living representatives.

Moreover, the blanks between strata in any particular vertical series are amply filled in with records in the form of thick strata in some geographically distant series. For example, in some regions Silurian rocks are directly overlaid by the coal measures; but elsewhere this sudden break is filled in with the Devonian rocks that tell of a great "age of fishes." So commonly are breaks in the strata in one region filled up in another that we are forced to conclude that the record shown by any single vertical series is of but local significance--telling, perhaps, of a time when that particular sea-bed oscillated above the water-line, and so ceased to receive sediment until some future age when it had oscillated back again. But if this be the real significance of the seemingly sudden change from stratum to stratum, then the whole case for catastrophism is hopelessly lost; for such breaks in the strata furnish the only suggestion geology can offer of sudden and catastrophic changes of wide extent.

Let us see how Lyell elaborates these ideas, particularly with reference to the rotation of species.(2)

"I have deduced as a corollary," he says, "that the species existing at any particular period must, in the course of ages, become extinct, one after the other. 'They must die out,' to borrow an emphatic expression from Buffon, 'because Time fights against them.' If the views which I have taken are just, there will be no difficulty in explaining why the habitations of so many species are now restrained within exceeding narrow limits. Every local revolution tends to circumscribe the range of some species, while it enlarges that of others; and if we are led to infer that new species originate in one spot only, each must require time to diffuse itself over a wide area. It will follow, therefore, from the adoption of our hypothesis that the recent origin of some species and the high antiquity of others are equally consistent with the general fact of their limited distribution, some being local because they have not existed long enough to admit of their wide dissemination; others, because circumstances in the animate or inanimate world have occurred to restrict the range within which they may once have obtained....

"If the reader should infer, from the facts laid before him, that the successive extinction of animals and plants may be part of the constant and regular course of nature, he will naturally inquire whether there are any means provided for the repair of these losses? Is it possible as a part of the economy of our system that the habitable globe should to a certain extent become depopulated, both in the ocean and on the land, or that the variety of species should diminish until some new era arrives when a new and extraordinary effort of creative energy is to be displayed? Or is it possible that new species can be called into being from time to time, and yet that so astonishing a phenomenon can escape the naturalist?

"In the first place, it is obviously more easy to prove that a species once numerously represented in a given district has ceased to be than that some other which did not pre-exist had made its appearance--assuming always, for reasons before stated, that single stocks only of each animal and plant are originally created, and that individuals of new species did not suddenly start up in many different places at once.

"So imperfect has the science of natural history remained down to our own times that, within the memory of persons now living, the numbers of known animals and plants have doubled, or even quadrupled, in many classes. New and often conspicuous species are annually discovered in parts of the old continent long inhabited by the most civilized nations. Conscious, therefore, of the limited extent of our information, we always infer, when such discoveries are made, that the beings in question bad previously eluded our research, or had at least existed elsewhere, and only migrated at a recent period into the territories where we now find them.

"What kind of proofs, therefore, could we reasonably expect to find of the origin at a particular period of a new species?

"Perhaps, it may be said in reply, that within the last two or three centuries some forest tree or new quadruped might have been observed to appear suddenly in those parts of England or France which had been most thoroughly investigated--that naturalists might have been able to show that no such being inhabited any other region of the globe, and that there was no tradition of anything similar having been observed in the district where it had made its appearance.

"Now, although this objection may seem plausible, yet its force will be found to depend entirely on the rate of fluctuation which we suppose to prevail in the animal world, and on the proportions which such conspicuous subjects of the animal and vegetable kingdoms bear to those which are less known and escape our observation. There are perhaps more than a million species of plants and animals, exclusive of the microscopic and infusory animalcules, now inhabiting the terraqueous globe, so that if only one of these were to become extinct annually, and one new one were to be every year called into being, much more than a million of years might be required to bring about a complete revolution of organic life.

"I am not hazarding at present any hypothesis as to the probable rate of change, but none will deny that when the annual birth and the annual death of one species on the globe is proposed as a mere speculation, this, at least, is to imagine no slight degree of instability in the animate creation. If we divide the surface of the earth into twenty regions of equal area, one of these might comprehend a space of land and water about equal in dimensions to Europe, and might contain a twentieth part of the million of species which may be assumed to exist in the animal kingdom. In this region one species only could, according to the rate of mortality before assumed, perish in twenty years, or only five out of fifty thousand in the course of a century. But as a considerable portion of the whole world belongs to the aquatic classes, with which we have a very imperfect acquaintance, we must exclude them from our consideration, and, if they constitute half of the entire number, then one species only might be lost in forty years among the terrestrial tribes. Now the mammalia, whether terrestrial or aquatic, bear so small a proportion to other classes of animals, forming less, perhaps, than a thousandth part of a whole, that, if the longevity of species in the different orders were equal, a vast period must elapse before it would come to the turn of this conspicuous class to lose one of their number. If one species only of the whole animal kingdom died out in forty years, no more than one mammifer might disappear in forty thousand years, in a region of the dimensions of Europe.

"It is easy, therefore, to see that in a small portion of such an area, in countries, for example, of the size of England and France, periods of much greater duration must elapse before it would be possible to authenticate the first appearance of one of the larger plants or animals, assuming the annual birth and death of one species to be the rate of vicissitude in the animal creation throughout the world."(3)

In a word, then, said Lyell, it becomes clear that the numberless species that have been exterminated in the past have died out one by one, just as individuals of a species die, not in vast shoals; if whole populations have passed away, it has been not by instantaneous extermination, but by the elimination of a species now here, now there, much as one generation succeeds another in the life history of any single species. The causes which have brought about such gradual exterminations, and in the long lapse of ages have resulted in rotations of population, are the same natural causes that are still in operation. Species have died out in the past as they are dying out in the present, under influence of changed surroundings, such as altered climate, or the migration into their territory of more masterful species. Past and present causes are one--natural law is changeless and eternal.

Such was the essence of the Huttonian doctrine, which Lyell adopted and extended, and with which his name will always be associated. Largely through his efforts, though of course not without the aid of many other workers after a time, this idea--the doctrine of uniformitarianism, it came to be called--became the accepted dogma of the geologic world not long after the middle of the nineteenth century. The catastrophists, after clinging madly to their phantom for a generation, at last capitulated without terms: the old heresy became the new orthodoxy, and the way was paved for a fresh controversy.

THE ORIGIN OF SPECIES

The fresh controversy followed quite as a matter of course. For the idea of catastrophism had not concerned the destruction of species merely, but their introduction as well. If whole faunas had been extirpated suddenly, new faunas had presumably been introduced with equal suddenness by special creation; but if species die out gradually, the introduction of new species may be presumed to be correspondingly gradual. Then may not the new species of a later geological epoch be the modified lineal descendants of the extinct population of an earlier epoch?

The idea that such might be the case was not new. It had been suggested when fossils first began to attract conspicuous attention; and such sagacious thinkers as Buffon and Kant and Goethe and Erasmus Darwin had been disposed to accept it in the closing days of the eighteenth century. Then, in 1809, it had been contended for by one of the early workers in systematic paleontology--Jean Baptiste Lamarck, who had studied the fossil shells about Paris while Cuvier studied the vertebrates, and who had been led by these studies to conclude that there had been not merely a rotation but a progression of life on the globe. He found the fossil shells--the fossils of invertebrates, as he himself had christened them--in deeper strata than Cuvier's vertebrates; and he believed that there had been long ages when no higher forms than these were in existence, and that in successive ages fishes, and then reptiles, had been the highest of animate creatures, before mammals, including man, appeared. Looking beyond the pale of his bare facts, as genius sometimes will, he had insisted that these progressive populations had developed one from another, under influence of changed surroundings, in unbroken series.

Of course such a thought as this was hopelessly misplaced in a generation that doubted the existence of extinct species, and hardly less so in the generation that accepted catastrophism; but it had been kept alive by here and there an advocate like Geoffrey Saint-Hilaire, and now the banishment of catastrophism opened the way for its more respectful consideration. Respectful consideration was given it by Lyell in each recurring edition of his Principles, but such consideration led to its unqualified rejection. In its place Lyell put forward a modified hypothesis of special creation. He assumed that from time to time, as the extirpation of a species had left room, so to speak, for a new species, such new species had been created de novo; and he supposed that such intermittent, spasmodic impulses of creation manifest themselves nowadays quite as frequently as at any time in the past. He did not say in so many words that no one need be surprised to-day were he to see a new species of deer, for example, come up out of the ground before him, "pawing to get free," like Milton's lion, but his theory implied as much. And that theory, let it be noted, was not the theory of Lyell alone, but of nearly all his associates in the geologic world. There is perhaps no other fact that will bring home to one so vividly the advance in thought of our own generation as the recollection that so crude, so almost unthinkable a conception could have been the current doctrine of science less than half a century ago.

This theory of special creation, moreover, excluded the current doctrine of uniformitarianism as night excludes day, though most thinkers of the time did not seem to be aware of the incompatibility of the two ideas. It may be doubted whether even Lyell himself fully realized it. If he did, he saw no escape from the dilemma, for it seemed to him that the record in the rocks clearly disproved the alternative Lamarckian hypothesis. And almost with one accord the paleontologists of the time sustained the verdict. Owen, Agassiz, Falconer, Barrande, Pictet, Forbes, repudiated the idea as unqualifiedly as their great predecessor Cuvier had done in the earlier generation. Some of them did, indeed, come to believe that there is evidence of a progressive development of life in the successive ages, but no such graded series of fossils had been discovered as would give countenance to the idea that one species had ever been transformed into another. And to nearly every one this objection seemed insuperable.

But in 1859 appeared a book which, though not dealing primarily with paleontology, yet contained a chapter that revealed the geological record in an altogether new light. The book was Charles Darwin's Origin of Species, the chapter that wonderful citation of the "Imperfections of the Geological Record." In this epoch-making chapter Darwin shows what conditions must prevail in any given place in order that fossils shall be formed, how unusual such conditions are, and how probable it is that fossils once imbedded in sediment of a sea-bed will be destroyed by metamorphosis of the rocks, or by denudation when the strata are raised above the water-level. Add to this the fact that only small territories of the earth have been explored geologically, he says, and it becomes clear that the paleontological record as we now possess it shows but a mere fragment of the past history of organisms on the earth. It is a history "imperfectly kept and written in a changing dialect. Of this history we possess the last volume alone, relating only to two or three countries. Of this volume only here and there a short chapter has been preserved, and of each page only here and there a few lines." For a paleontologist to dogmatize from such a record would be as rash, he thinks, as "for a naturalist to land for five minutes on a barren point of Australia and then discuss the number and range of its productions."

This citation of observations, which when once pointed out seemed almost self-evident, came as a revelation to the geological world. In the clarified view now possible old facts took on a new meaning. It was recalled that Cuvier had been obliged to establish a new order for some of the first fossil creatures he examined, and that Buckland had noted that the nondescript forms were intermediate in structure between allied existing orders. More recently such intermediate forms had been discovered over and over; so that, to name but one example, Owen had been able, with the aid of extinct species, to "dissolve by gradations the apparently wide interval between the pig and the camel." Owen, moreover, had been led to speak repeatedly of the "generalized forms" of extinct animals, and Agassiz had called them "synthetic or prophetic types," these terms clearly implying "that such forms are in fact intermediate or connecting links." Darwin himself had shown some years before that the fossil animals of any continent are closely related to the existing animals of that continent--edentates predominating, for example, in South America, and marsupials in Australia. Many observers had noted that recent strata everywhere show a fossil fauna more nearly like the existing one than do more ancient strata; and that fossils from any two consecutive strata are far more closely related to each other than are the fossils of two remote formations, the fauna of each geological formation being, indeed, in a wide view, intermediate between preceding and succeeding faunas.

So suggestive were all these observations that Lyell, the admitted leader of the geological world, after reading Darwin's citations, felt able to drop his own crass explanation of the introduction of species and adopt the transmutation hypothesis, thus rounding out the doctrine of uniformitarianism to the full proportions in which Lamarck had conceived it half a century before. Not all paleontologists could follow him at once, of course; the proof was not yet sufficiently demonstrative for that; but all were shaken in the seeming security of their former position, which is always a necessary stage in the progress of thought. And popular interest in the matter was raised to white heat in a twinkling.

So, for the third time in this first century of its existence, paleontology was called upon to play a leading role in a controversy whose interest extended far beyond the bounds of staid truth-seeking science. And the controversy waged over the age of the earth had not been more bitter, that over catastrophism not more acrimonious, than that which now raged over the question of the transmutation of species. The question had implications far beyond the bounds of paleontology, of course. The main evidence yet presented had been drawn from quite other fields, but by common consent the record in the rocks might furnish a crucial test of the truth or falsity of the hypothesis. "He who rejects this view of the imperfections of the geological record," said Darwin, "will rightly reject the whole theory."

With something more than mere scientific zeal, therefore, paleontologists turned anew to the records in the rocks, to inquire what evidence in proof or refutation might be found in unread pages of the "great stone book." And, as might have been expected, many minds being thus prepared to receive new evidence, such evidence was not long withheld.

FOSSIL MAN

Indeed, at the moment of Darwin's writing a new and very instructive chapter of the geologic record was being presented to the public--a chapter which for the first time brought man into the story. In 1859 Dr. Falconer, the distinguished British paleontologist, made a visit to Abbeville, in the valley of the Somme, incited by reports that for a decade before bad been sent out from there by M. Boucher de Perthes. These reports had to do with the alleged finding of flint implements, clearly the work of man, in undisturbed gravel-beds, in the midst of fossil remains of the mammoth and other extinct animals. What Falconer saw there and what came of his visit may best be told in his own words:

"In September of 1856 I made the acquaintance of my distinguished friend M. Boucher de Perthes," wrote Dr. Falconer, "on the introduction of M. Desnoyers at Paris, when he presented to me the earlier volume of his Antiquites celtiques, etc., with which I thus became acquainted for the first time. I was then fresh from the examination of the Indian fossil remains of the valley of the Jumna; and the antiquity of the human race being a subject of interest to both, we conversed freely about it, each from a different point of view. M. de Perthes invited me to visit Abbeville, in order to examine his antediluvian collection, fossil and geological, gleaned from the valley of the Somme. This I was unable to accomplish then, but I reserved it for a future occasion.

"In October, 1856, having determined to proceed to Sicily, I arranged by correspondence with M. Boucher de Perthes to visit Abbeville on my journey through France. I was at the time in constant communication with Mr. Prestwich about the proofs of the antiquity of the human race yielded by the Broxham Cave, in which he took a lively interest; and I engaged to communicate to him the opinions at which I should arrive, after my examination of the Abbeville collection. M. de Perthes gave me the freest access to his materials, with unreserved explanations of all the facts of the case that had come under his observation; and having considered his Menchecourt Section, taken with such scrupulous care, and identified the molars of elephas primigenius, which he had exhumed with his own hands deep in that section, along with flint weapons, presenting the same character as some of those found in the Broxham Cave, I arrived at the conviction that they were of contemporaneous age, although I was not prepared to go along with M. de Perthes in all his inferences regarding the hieroglyphics and in an industrial interpretation of the various other objects which he had met with."(4)

That Dr. Falconer was much impressed by the collection of M. de Perthes is shown in a communication which he sent at once to his friend Prestwich:

"I have been richly rewarded," he exclaims. "His collection of wrought flint implements, and of the objects of every description associated with them, far exceeds everything I expected to have seen, especially from a single locality. He has made great additions, since the publication of his first volume, in the second, which I now have by me. He showed me flint hatchets which HE HAD DUG UP with his own hands, mixed INDISCRIMINATELY with molars of elephas primigenius. I examined and identified plates of the molars and the flint objects which were got along with them. Abbeville is an out-of-the-way place, very little visited; and the French savants who meet him in Paris laugh at Monsieur de Perthes and his researches. But after devoting the greater part of a day to his vast collection, I am perfectly satisfied that there is a great deal of fair presumptive evidence in favor of many of his speculations regarding the remote antiquity of these industrial objects and their association with animals now extinct. M. Boucher's hotel is, from the ground floor to garret, a continued museum, filled with pictures, mediaeval art, and Gaulish antiquities, including antediluvian flint-knives, fossil-bones, etc. If, during next summer, you should happen to be paying a visit to France, let me strongly recommend you to come to Abbeville. I am sure you would be richly rewarded."(5)