Chapter II: Preface (2)
Frustule cylindrical, less in length than the diameter, in zone view undulated. Valve divided into six or more sectors alternately raised and depressed, areolate and punctate, varying in the alternate divisions. The areolation is confined to the outer layer of the valve while the punctation is usually on an inner valve often found detached. Processes on the border, three or more. Umbilicus circular or angular, hyaline.
ANALYSIS OF SPECIES
Sectors, six undulatus
Sectors, eight or more, cellular heliopelta
Sectors, fourteen, punctate vulgaris
ACTINOPTYCHUS UNDULATUS (KUETZ.) RALFS
Valve areolate and punctate in quincunx, divided into six equal sectors, alternately elevated or depressed, their areolations appearing different. Margin well defined. Umbilicus smooth, hexagonal. Processes three, sometimes six, inserted within the margin of each alternate division. Very variable in size and appearance.
This is the Actinocyclus of Bailey, figured and described in Amer. Jour. Science, 1842, p. 93, Pl. 2, Fig. 11, but not named. Kuetzing describes and names it and refers to Bailey.
_Actinoptychus omphalopelta_ Ehr.
_Actinoptychus cellulosa_ Ehr., H. L. Smith Sp. Typ., 384.
Quite common in marine and brackish water and in the blue clay.
Pl. 4, Figs. 1, 2, 4 and 6.
ACTINOPTYCHUS VULGARIS VAR. INTERRUPTA N. VAR.
Valve with fourteen sectors, the alternate ones divided by a smooth lanceolate space for about one-half the radius, forming with the smooth, circular umbilicus a seven pointed star. The sectors thus divided have coarser puncta in quincunx than the other sectors, ending in a smooth area near the margin, and also larger black puncta scattered from the centre to the semi-radius.
Near A. vulgaris var. neogradensis Pant.
Blue clay. Not common.
Pl. 4, Fig. 5.
{25}ACTINOPTYCHUS HELIOPELTA GRUN. VAR.?
Valve circular, sectors, eight, umbilicus circular, without rays; border wide, cellular, with distinct rays. Inserted at a distance within the inner edge of the border are large processes, one on each of four alternate sectors, and two on each of the others. The sectors are cellulate and punctate.
Near A. heliopelta var. versicolor Brun., which, however, in the specimen in my collection from Atlantic City (artesian well), has a greater number of processes and they are situated on the edge of the border.
Outcrop at Buckshutem, N. J. Rare.
Pl. 4, Fig. 3.
It has been quite well determined, I think, that the typical forms of A. heliopelta occur at the base of the Miocene. At Rock Hall, Md., on the eastern shore of Chesapeake Bay, at a depth of from 21 to 130 ft., and at Wildwood, N. J., at a depth of from 78 to 179 ft., diatomaceous beds occur considered by Mr. Lewis Woolman (Geol. Surv. of N. J., 1898, pp. 116-121) "as synchronous in age," the former being deposited in the Delaware River Delta and the latter in the Chesapeake in post-miocene times. In each of these beds a small form of A. heliopelta is rarely found. The material at Buckshutem is post-miocene, and the form here figured shows a marked variation from the Miocene species and a gradual approach toward A. undulatus.
POLYMYXUS L. W. BAIL. (1855)
Valve circular, usually divided into fourteen sectors which are on the same plane at the centre, but the alternate ones are elevated into mammillated projections terminated by small processes on the margin. Zone view rectangular with undulations subconical, terminated by the processes.
POLYMYXUS CORONALIS L. W. BAIL.
Central space hyaline, rounded or slightly stellate, from which radiate rows of fine puncta in quincunx, shown in the figure only on the alternate elevations, the depressed interspaces being out of focus. The mammillæ are stated by Bailey to vary from six to ten.
Very rare in the blue clay (Walnut St. Bridge). Occurs also in the Wildwood deposit (Bull. Torrey Bot. Club, 1895, p. 261).
Pl. 4, Fig. 7, and Pl. 5, Fig. 2.
3. EUPODISCEÆ
_Aulacodiscinæ._--Valves with mammiform elevations near the border surmounted by nipple-like processes.
AULACODISCUS--THE ONLY GENUS AS ABOVE
_Eupodiscinæ._--Valves with ocelli.
(1) _Actinocyclus._--Valve with one small ocellus; striæ radial.
(2) _Eupodiscus._--Valve with one or more ocelli; striæ not radial.
(3) _Auliscus._--Valve with large, elevated ocelli. Central area hyaline. Markings granular and costate.
(4) _Pseudauliscus._--Valve with radiating granules. No central space.
{26}AULACODISCUS EHR. (1844) em. RATTR. (1888)
(aulax, a furrow, and discus)
Valve usually circular, plane or with an elevated zone, frequently inflated beneath the processes; central space irregular or rounded, sometimes absent; markings granular, radial, sometimes in a reticulum.
The genus comprises more than one hundred species most of which are fossil, and is represented in this locality by a single form, _A. argus_, included by Rattray in his section "Retiformes," distinguished by the presence of a reticulum.
AULACODISCUS ARGUS (EHR.) SCHMIDT
Frustule in zone view elliptical. Valve circular, 125-190 µ in diam., closely covered with two kinds of markings, one, a mesh of large, radiating, angular cells, the outer plate, and the other, radiating rows of circular granules with hyaline spaces intervening and closer near the border, forming the inner plate which can occasionally be seen detached. Central space absent. The walls of the angular cells are crossed with fine lines and are probably composed of granules compressed so closely as to produce partial opacity, the depth of which depends in a measure not only on the superposition of the two plates, but on the relative closeness and thickness of the cell-walls. In a fully-developed specimen the effect is to produce more or less triangular cells containing three or four granules. In some cases the opacity is so great as to render detail invisible.
In the figure the valve is supposed to be divided into three sectors, illustrating at "a" the lower plate, at "c" the combination of the upper and lower plates, and in the other sector the cellular mesh of the upper plate. Processes, usually three, quite robust and inserted at from one-fourth to one-fifth the length of the radius from the border which is striated on the inner side. A form with four processes is found in the lower blue clay.
_Tripodiscus argus_ Ehr.
_Eupodiscus argus_ (Ehr.) Wm. Sm.
Not uncommon in the blue clay.
Pl. 4, Fig. 8.
ACTINOCYCLUS EHR. (1837)
(actis, a ray, and cyclos)
Valve circular or elliptical; surface flat at the centre, sloping toward the border. Central space usually evident, rounded or irregular. Markings rounded, granular, punctiform, in radial, or nearly radial, rows, sometimes fasciculate. A nodule, more or less evident, is found near the border which is usually striate.
Chromatophores round discs or granules.
ANALYSIS OF SPECIES
Valve circular, rows radial, hyaline lines at the border barkleyi
Valve circular, rows fasciculate moniliformis
Valve elliptical ellipticus
The nodule is generally supposed to be a thickening of the cell-wall, and, in the opinion of Rattray, a projection outward, but "whether there may not be at the same time a slight inward protuberance is difficult to determine," though, as a rule, he seems to "think there is not."
{27}ACTINOCYCLUS BARKLEYI VAR. AGGREGATA RATTR.
Surface flat from centre to semi-radius. Central space irregular, sometimes with a few scattered granules. Markings round with central dots distinct, about 7 at the centre, decreasing in straight radial rows to 12 in 10 µ at the border, where they form moniliform striæ. Border narrow with striæ about 16 in 10 µ. Hyaline interspaces at the origin of the shorter rows, but not at equal intervals. At the border, linear hyaline spaces occur at somewhat irregular intervals between the moniliform striæ owing to the termination of certain radial rows before they reach the circumference. Nodule small, from one-seventh to one-fourth the radius from the border.
According to Rattray the distinction between A. ralfsii and A. barkleyi is partly in the absence of the zone arrangement of the hyaline spaces in the latter, and to the slight differences in the number of granules. The variety aggregata differs from the type form of barkleyi mainly in the distance of the nodule from the border. I have specimens from the blue clay material at Walnut St. Bridge, and from Smith's Island, in which the distance from the border in one case is, as stated above, quite different from that in the other. In specimens from Morris Cove, Conn., the locality referred to by Rattray, variations occur.
Blue clay.
Pl. 6, Fig. 1.
In the figure the subulate hyaline spaces at the border are, in some instances, wider than usual.
ACTINOCYCLUS MONILIFORMIS RALFS
Surface flat, from centre to about five-sixths of the radius. Central space rounded, with one or more granules. Markings, 8 in 10 µ, round, in radial rows, fasciculate, the oblique transverse rows irregular, very slightly decreasing until near the edge of the flattened zone, and then suddenly decreasing and appearing as decussating lines oblique to the border. Apiculi distinct, interfasciculate within the border. Nodule quite evident, surrounded by a rather wide irregular hyaline space on the margin of the flattened zone in the middle of the fasciculus. Border wide, with striæ about 20 in 10 µ.
Blue clay. Port Penn. Not common.
Pl. 6, Fig. 2.
Equivalent to Actinocyclus ehrenbergii, H. L. S. Type Slide 10.
In a valve from Port Penn, Delaware Bay, two nodules occur nearly opposite each other.
ACTINOCYCLUS ELLIPTICUS VAR. DELAWARENSIS N. VAR.
Valve rhombic-elliptical. Markings somewhat angular, 6 in 10 µ at the centre where they are sub-concentric, thence decreasing in lines radiating more or less toward the border, where they suddenly become punctiform, striæ about 20 in 10 µ. Border equal to one-fifth the radius. A nodule is found on the inner side of the border. Apiculi apparently absent.
The markings are larger than in the Richmond forms which are associated by Rattray with Actinocyclus ellipticus Grun. The form corresponds closely to Witt's Cestodiscus ovalis var.? (Witt, Polierschief. von Archangelsk-Kurojedowo, Pl. 8, Fig. 2), except as to the border. It does not answer to Van Heurck's figure or any other.
Blue clay. Very rare.
Pl. 3, Fig. 6.
{28}EUPODISCUS EHR. (1844)
(eu, well, pous, a foot, and discus)
Valve circular, 45-117 µ in diam. (De Toni). Central space absent, surface plane with angular cells. At the border short, circular processes or ocelli.
EUPODISCUS RADIATUS BAIL.
Valve with radiating hexagonal cells, sometimes slightly curved toward the large ocelli inserted near the border which are hyaline at the centre. Border wide, coarsely striate.
The number of ocelli heretofore recorded is four. Specimens with five processes are found in the artesian well at St. Augustine, Fla., and in material at Twelfth and Brandywine Sts. Mr. Hugo Bilgram has discovered valves with three and six ocelli.
Not common in the blue clay, but abundant along the southern coast of the Atlantic states and the Gulf of Mexico.
Not Eupodiscus radiatus Wm. Sm, which is Biddulphia smithii (Ralfs) V. H.
Pl. 5, Fig. 3.
AULISCUS EHR. (1843)
(aulax, a furrow, referring to the grooves in certain species, according to De Toni, but preferably from auliscos, a small reed, referring to the processes?)
Frustule cylindrical; zone with longitudinal rows of fine puncta. Valve circular or elliptical, plane except near the processes; central area hyaline, usually circular. Markings of two kinds, granules radiating or scattered and radiating, costate lines, prominent or indistinct. Processes, two or three, large, short, cylindrical, with hyaline surface, near the ends of the major axis in a line oblique to it.
Auliscus is divided by Rattray into fourteen sections, defined chiefly by the character and arrangement of the markings. About eighty species are described, but as many of the forms are fossil, occuring in the Miocene of California, Oamaru and elsewhere, and as so few species are found in this locality, I shall refer but briefly to this division.
_Striolati._--No transverse median areas, striæ
inconspicuous punctatus
_Lineolati._--Markings distinct, pruinose, interrupted pruinosus
_Costati._--Transverse median areas usually distinct, sculptus
markings continuous, costate cælatus
AULISCUS PUNCTATUS BAIL.
Valve broadly elliptical, or suborbicular, covered with delicate interrupted striæ radiating in sinuous lines to the circumference, more evident on the transverse median area; puncta 3 in 10 µ, grouped into a rounded area on each side of the median line, elsewhere scattered. Central space rounded, processes two, large, suborbicular.
Port Penn, Delaware River. Rare.
Pl. 5, Fig. 6.
AULISCUS PRUINOSUS BAIL.
Valve elliptical, with distinct, interrupted, pruinose, irregular markings diverging in curved lines toward the circumference in the median part and converging toward the processes, interspersed with numerous darker markings having the appearance of apiculi. Central space nearly circular, sometimes with several granules. Processes large near the ends of the major axis and not oblique to it, or scarcely so, the edges with a crenulate border.
Blue clay. Rather rare.
Pl. 5, Fig. 8.
{29}AULISCUS SCULPTUS (WM. SM.) RALFS
Valve elliptical or subcircular, median areas distinct, rounded, circumscribed by coarse distant costæ radiating near the border where they are more evident, and converging toward the processes. Central space rounded, sometimes indefinite. Processes, two, circular.
Typical specimens show wide, coarse, distant costæ, but, in some cases, the median areas are indistinctly outlined.
Blue clay.
Pl. 5, Fig. 5.
AULISCUS CÆLATUS BAIL.
Valve elliptical or subcircular, with radiating costæ, more evident around the median areas and at the border, converging toward the processes, with intermediate punctate radiating lines. Central space rounded or irregular. Processes circular.
A. sculptus has coarser costæ and the interspaces are hyaline, or apparently so, while in A. cælatus the punctate striæ between the costæ are more evident.
Blue clay. Not uncommon.
Pl. 5, Fig. 4.
Fig. 7 is a small, indefinite form intermediate between A. sculptus and A. cælatus. The numerous variations in this genus make it difficult to satisfactorily differentiate the species. The size of the four above described varies from 40 to 150 µ.
PSEUDAULISCUS A. S. (1875) AND LEUDUGER-FORTMOREL (1879)
Valve circular or subcircular, nearly flat or depressed at the centre. Central space not evident. Processes circular, with narrow border, near the circumference. Border narrow, striated. Markings granular, radiating, sometimes interspersed with striæ and apiculi.
Differs from Auliscus chiefly in the absence of a central space and costæ.
PSEUDAULISCUS RADIATUS (BAIL.) RATTR.
Valve circular, or nearly so, flat. Central area with scattered granules radiating and increasing in size outward in diverging rows toward the border which is coarsely striated. Processes, two, circular. Two small apiculi are inserted at about one-fifth the radius from the border near the ends of the minor axis.
Blue clay. Rare.
Pl. 5, Fig. 9.
The apiculi are not always figured. They appear in a number of specimens from the Miocene of Maryland, Atlantic City, Harvey Cedars and Newbern.
PSEUDAULISCUS SPINOSUS (CHRISTIAN) RATTR.
Valve subcircular or slightly quadrangular, depressed at the centre and rising to an elevated zone near the border, the two zones separated by a distinct line. The inner zone indistinctly reticulate with fine puncta radiating from the centre and apiculi at intervals. The outer zone with smaller apiculi surrounding the inner zone and with intermingled rows of fine puncta and interrupted diverging striæ. Near each end of the minor axis is a rather long, robust spine inserted at one-fourth the radius from the border which is narrow and striated. Processes circular, close to the circumference.
_Auliscus spinosus_ Christian.
Blue clay. Rare.
Pl. 5, Fig. 10.
The genus is named by Schmidt, described by Leuduger-Fortmorel and emendated by Rattray.
{30}BIDDULPHIOIDEÆ
BIDDULPHIEÆ
(_a_) _Triceratiinæ._--Frustule cylindrical or prismatic, with three or more sides.
(_b_) _Biddulphiinæ._--Frustule cylindroid; valve with ends elevated into round processes or long horns.
(_c_) _Anauleæ._--Valve elliptical, lunate or triangular, with internal septa.
(_d_) _Euodieæ._--Frustule cuneate in zone view; valve lunate.
(_a_) TRICERATIINÆ
(1) _Ditylum._--Frustule imperfectly siliceous. Zone with numerous divisions. Valve with central spine.
(2) _Trinacria._--Processes with sharp spines.
DITYLUM BAIL. (1861)
(dis, two, and tyle, a swelling, referring to the outline of the frustule)
Frustule quadrangular, convex at the ends. Valve triangular, with undulating sides, the angles ending in a sharp point surmounted by a bristle. Surface of valve convex at centre from which projects a long stout spine.
DITYLUM INTRICATUM (WEST) GRUN.
Valve with the angles separated from the central part by lines imitating septa. Surface with radiating lines of fine puncta.
Blue clay. Rare.
Pl. 6, Fig. 4.
Detached valves only have been found in the blue clay. The form is regarded as but slightly siliceous and, therefore, the zone or girdle not being found in the fossil deposits, I am unable to illustrate it from material in the vicinity. On Plate 38, Figs. 6 and 7, I have sketched the zone and valve views of specimens found recently at Vera Cruz and labelled by H. L. Smith Triceratum intricatum West. I can find no difference between the recent and fossil forms of the valves. The zone is covered with fine puncta in quincunx, not visible under ordinary illumination.
The form as figured in Plate 6 corresponds to the figure of Lithodesmium undulatum Ehr. in Van Heurck, and West, in describing the Triceratium undulatum Wm. Sm. (figured as T. striolatum), thought that his T. intricatum was distinct from Ehrenberg's form on the ground that the latter came from the "Bermuda" (Nottingham) earth and must be strongly siliceous. Lithodesmium is characterized by the envelopment of the frustules by a cellular membrane which does not appear, evidently, in Ditylum. D. brightwellii is distinguished by its crown of spines on the margin; otherwise it closely resembles D. intricatum.
TRINACRIA HEIB. (1863)
(treis, three, and acra, a point)
Valve triangular, angles elevated into spines. Cells at the margin large.
TRINACRIA PILEOLUS (EHR.) GRUN.
Valve with concave sides. Surface concave with unequal punctiform and scattered markings with central dots. Cells at the margin large, rounded. At the angles, which vary in elevation, a few puncta are seen.
_Triceratium pileolus_ Ehr.
Blue clay. Rare.
Pl. 6, Fig. 9.
{31}(_b_) BIDDULPHIINÆ
BIDDULPHIA GRAY (1831) em. VAN HEURCK (1885)
(a genus, constituted from Conferva biddulphiana of the English Botany, named after a Miss Biddulph)
Frustule prismatic or subcylindrical, concatenate, filamentous, or in zig-zag, or, as usually found, free. Zone well developed. Valve triangular, polygonal, elliptic or subcircular, convex, more or less elevated at the angles into processes or horns. Markings cellular or punctate. Chromatophores, small plates of various forms.
KEY TO THE SPECIES
Valves costate biddulphiana
Valves not costate:
Markings cellular, angles elevated into horns favus
angles not elevated antediluviana
Markings punctate, angles with subconical processes and
long spines granulata
spines short rhombus
spines minute smithii
processes truncate, valve elliptical turgida
valve orbicular lævis
processes absent, valve divided by irregular lines alternans
not so divided reticulum
BIDDULPHIA BIDDULPHIANA (SMITH)
Frustule quadrangular with convex ends and rounded angles. Valve elliptical with undulated sides, divided by septa into three or more sections. Processes large, rounded, globular or subconical. Zone varying in width. Surface with rounded reticulations in longitudinal and transverse rows, except at the centre where they are concentric and smaller.
_Conferva biddulphiana_ Smith (English Botany, 1807, Pl. 1762, upper figures).
_Diatoma biddulphianum_ Ag.
_Biddulphia pulchella_ Gray.
Blue clay. Hoboken Tunnel. Along the coast.
Pl. 7, Figs. 1, 2, 3, and 4.
Quite variable in size and number of septate divisions. Fig. 3 is an unusual form with narrow zone, having but one row of large reticulations, evidently a young frustule.
BIDDULPHIA FAVUS (EHR.) V. H.
Frustule quadrangular, elevated at the angles into subconical processes oblique to the longitudinal axis. Valve triangular or quadrangular, plane, of two layers, the outer layer composed of large hexagonal cells in rows parallel to the sides, the inner of small puncta radiating from the centre. Zone punctate in quincunx, never found open.
_Triceratium favus_ Ehr.
Blue clay. Common along the coast.
The quadrangular form occurs only southward.
Pl. 6, Fig. 6. At "a" a cell showing the lower punctate layer. Pl. 40, Fig. 16, a transverse section of a portion of the valve showing the cellular structure and the punctated lower stratum.
{32}BIDDULPHIA ANTEDILUVIANA (EHR.) V. H.
Frustules quadrangular, sometimes united in zig-zag chains. Valve quadrangular with more or less concave sides, sometimes cruciform. Surface with angular cells arranged in concentric and radiating lines increasing toward the circumference. At each angle is a large, rounded process, which, as well as the secondary layer, scarcely visible, is finely punctate.
_Amphitetras antediluviana_ Ehr.
_Amphitetras tessellata_ Shad.
Blue clay. Rare.
Pl. 6, Fig. 3.
A cruciform variety occurs at Pensauken, N. J., artesian well (Coll. F. J. Keeley).
BIDDULPHIA GRANULATA ROPER
Valve elliptical-lanceolate, convex, with diagonal rows of puncta 12 in 10 µ and sometimes with small scattered spurs. Processes inflated at the base, obtuse at the ends, which are curved outward toward alternate sides. Near each process and on opposite sides of the longitudinal axis is placed a stout spine bent or curved inward near the middle. Connective zone with diagonal rows of puncta smaller than those on the valve.
Pavonia, N. J., artesian well. Fossil in the Pleistocene. Along the coast. Not common.
Pl. 7, Fig. 6.
BIDDULPHIA RHOMBUS (EHR.) WM. SM.
Valve rhomboidal, sometimes triangular, with subconical processes. Surface convex with hexagonal reticulations, 7-9 in 10 µ, irregular at the centre and radiating to the circumference. Minute spurs are scattered over the surface, and on each side are usually two or three short spines.
Common along the coast and fossil in the Miocene and later deposits.
Pl. 7, Fig. 5 (somewhat inclined, as usually seen).
BIDDULPHIA SMITHII (RALFS) V. H.
Valve orbicular, convex, with reticulations 5 in 10 µ radiating from the centre and decreasing toward the margin and processes which are truncate. A short spine is found on each side half way between the processes. Zone narrow with fine puncta 12 in 10 µ in longitudinal rows.
_Cerataulus smithii_ Ralfs.
_Eupodiscus radiatus_ Wm. Sm.
Blue clay. Along the coast southward.
Pl. 7, Fig. 8.
BIDDULPHIA TURGIDA (EHR.) WM. SM.
Valve elliptical or orbicular, surface convex. Processes very large, cylindrical, placed obliquely and inclined by the torsion of the frustule. Between the processes are two stout spines, one on each side, frequently forked at the ends. Puncta fine, irregular at the centre and radiating toward the circumference.
_Cerataulus turgidus_ Ehr.
Blue clay. Along the coast. Quite variable in size.
Pl. 7, Fig. 7.
{33}BIDDULPHIA LÆVIS EHR.
Valve suborbicular or triangular, with short, truncate processes. Surface with fine puncta about 13 in 10 µ radiating in straight or curved lines toward the circumference and with fine spurs at intervals. Nearer one process than the other, and about half way between centre and circumference, are two small spines, one on each side. Quite variable in size.
Blue clay. Common along the coast.
Pl. 7, Fig. 9.
Fig. 10 (magnification about 260 diameters only) illustrates sporangial frustules discovered by Mr. T. Chalkley Palmer at Reedy Island, Delaware River. In frustules having a cylindrical form, the endochrome lines the cell-walls in the form of granules which become congregated toward the centre in the sporangia.
BIDDULPHIA ALTERNANS (BAIL.) V. H.
Valve triangular or, rarely, quadrangular, with sides straight or slightly concave, usually unequal. Angles obtuse, separated from the centre by costate lines. Surface with puncta of irregular shape, large at the centre, with smaller puncta interspersed. In many valves several lines appearing like costæ extend inward from the border in various directions. Angles with small puncta in transverse and longitudinal rows.
_Triceratium alternans_ Bail.
Blue clay. Along the coast.
Pl. 6, Fig. 7 and probably Fig. 8.
BIDDULPHIA RETICULUM (EHR.)
Frustule quadrangular. Valve triangular with straight or concave sides and rounded angles. Surface convex at the centre and angles. Markings of unequal size, mostly larger at the centre, scattered; at the angles, small puncta in longitudinal rows.
_Triceratium sculptum_ Shad.
_Triceratium punctatum_ Br.
_Triceratium obtusum_ Br.
For explanation of the synonymy see "Biddulphoid Forms of N. A. Diat.," Proc. Acad. Nat. Sci., 1900, p. 724.
Blue clay. Along the coast.
Pl. 6, Fig. 5.
(_c_) ANAULEÆ
EUNOTOGRAMMA WEISSE (1854)
(eu, well, noton, a back, and gramma)
Frustule quadrangular. Valve elliptical or lunate divided by septa which constrict the margin. Surface flat with punctate markings.
EUNOTOGRAMMA LÆVE GRUN.
Valve lunate with obtuse ends. Septa, from four to eleven or more. Surface with puncta in transverse and longitudinal rows, sometimes indistinct and scattered.
Shark River. Rare. More common southward. Fossil at Buckshutem, N. J.
Pl. 7, Fig. 11, and Pl. 10, Fig. 15.
I am unable to distinguish between E. læve and E. debile, as intermediate forms occur.
{34}TERPSINOË EHR.
(terpsinoos, gladdening?)
Frustules quadrangular, adnate in filaments, usually free. Valve elliptical or triangular, with undulating sides divided by septa into three or more sections.
TERPSINOË AMERICANA (BAIL.) RALFS
Valve lobed at each end or angle. Central space rounded, hyaline. Surface with fine puncta in radiating lines.
Blue clay. Not common.
Pl. 6, Fig. 10.
TERPSINOË NOVÆ-CÆSAREÆ BOYER
Valve triangular, with concave sides and broad angles equally three-lobed, separated from the central part by septa. Central space small or absent. Puncta delicate, radiating or scattered. L. of side 62 µ.
Pleistocene clay at Buckshutem, N. J. Fossil at Wildwood, N. J.
T. americana, forma trigona Pant.? (Le Diatomiste, Vol. 2, p. 207.)
Pl. 6, Fig. 11.
(_d_) EUODIEÆ
EUODIA BAIL. (1860)
(derivation uncertain; apparently from euodia, fragrant, probably a euphemism)
Frustule in zone view cuneate. Valve semi-lunate, coscinodiscoid.
EUODIA GIBBA BAIL.
Valve with rounded markings, larger and scattered at the centre, radiating at the circumference and in indefinite straight rows at the semi-radius.
Delaware Bay (Mann).
Pl. 5, Fig. 1.
I have not seen this in the Philadelphia material. The figure is drawn from a specimen from the Gulf Stream, S. Atlantic.
{35}PENNATÆ
Valve zygomorphous. Structure pinnate, not concentric. Valve divided either by a true raphe or cleft or by a linear space or line imitating a raphe.
Divided into three Groups:
1. _Fragilarioideæ._--Valves without a raphe; usually with a pseudoraphe or median line.
2. _Naviculoideæ._--Either one or both valves with a true raphe.
3. _Surirelloideæ._--Valves in which the raphe is concealed near the margin on one or both sides of each valve in a more or less elevated keel or wing.
FRAGILARIOIDEÆ
(_a_) _Tabellarieæ._--Valve symmetrical with respect to both the longitudinal and transverse axes; septate, not cuneate.
(_b_) _Meridioneæ._--Valve symmetrical with respect to the longitudinal axis, asymmetrical to the transverse axis, cuneate, finely striated.
(_c_) _Fragilarieæ._--Valve of varied shape, not cuneate; costate or with transverse rows of puncta.
(_a_) TABELLARIEÆ
Frustule in zone view rectangular, in valve view linear or linear-elliptical, sometimes constricted in the middle, symmetrical to both axes, not cuneate; with two or more septa or annuli.
Chromatophores numerous, granular.
_Rhabdonema._--Frustules with numerous septate partitions having one or several foramina. Transverse costæ or rows of coarse puncta.
_Tabellaria._--Frustules with two to six nearly straight septa. Transverse striæ subtly punctate.
_Grammatophora._--Frustules with two sinuate perforate curved septa. Transverse striæ subtly punctate.
_Striatella._--Frustules with alternate partitions, septate or partly so.
_Attheya._--Frustules not septate but with numerous annuli.
RHABDONEMA KUETZ. (1844)
(rhabdos, a rod, and nema, a thread)
Frustules quadrangular, concatenate, composed of numerous septate partitions with transverse costæ or rows of puncta. Valves elliptical, with a pseudoraphe and transverse apparent costæ and punctate lines; the partitions with one or several foramina.
Chromatophores in rosettes of various kinds (Karsten); usually parallel to the septa.
RHABDONEMA ARCUATUM (LYNG.) KUETZ.
Valve hyaline at the ends, with transverse rows of puncta producing the appearance of costæ between the rows; pseudoraphe distinct; foramen single.
_Diatoma arcuatum_ Lyngbye.
Common along the coast.
Pl. 8, Figs. 1, 2, and 3; Pl. 40, Fig. 10.
{36}According to T. H. Buffham (Jour. Quek. M. C., Series 2, Vol. 2, p. 131), the frustules are of two kinds, those in which the length and breadth are the same and those which are much lengthened, with a wide hyaline girdle frequently in the middle. At the time of fructification the smaller frustules are attached to a larger one which produces a sporangium at the end of the girdle from which the other end of the frustule has disappeared, or, if the two halves of the frustule remain, two sporangia are formed.
RHABDONEMA MINUTUM KUETZ.
Frustules small; valve not smooth at the ends, elliptical or lanceolate-elliptical, with transverse rows of puncta; pseudoraphe distinct. Foramen single, alternating above and below in adjoining partitions.
Common in the blue clay and along the coast.
Pl. 8, Fig. 7 and Pl. 38, Fig. 11.
RHABDONEMA ADRIATICUM KUETZ.
Valve linear-lanceolate, with smooth angles; rows of puncta transverse, the intervals appearing as costæ, as in arcuatum. Foramina, three.
Blue clay in the Pensauken and Pavonia deposits and along the coast.
Pl. 8, Figs. 4, 5 and 6.
TABELLARIA EHR. (1839)
(tabella, a tablet)
Frustules quadrangular, adnate in filaments, frequently found in zig-zag chains, united by a gelatinous isthmus, at length separating. Valve linear, inflated in the middle and at the ends; striæ transverse.
Chromatophores numerous, small, along the zones.
TABELLARIA FENESTRATA (LYNG.) KUETZ.
Valve elongated; pseudoraphe narrow; transverse striæ faint. In the zone view a straight septum is shown at each end of a valve.
Common, especially in the cedar swamps and ponds of the Pine Barren region, N. J.
Pl. 8, Figs. 11 and 12.
TABELLARIA FLOCCULOSA (ROTH) KUETZ.
Valve linear, with median inflation larger than the terminal; pseudoraphe rather broad in the middle; transverse striæ subtly punctate. In zone view the frustules are quadrangular, or nearly so, with about six sometimes curved septa at one end alternating with those on the other end.
_Conferva flocculosa_ Roth.
Common especially in the Pine Barrens of New Jersey.
Pl. 8, Figs. 8, 9 and 10.
GRAMMATOPHORA EHR. (1840)
(from gramma, a letter, and phoreo, I bear)
Frustules quadrangular, adnate, in zig-zag, united by an isthmus, or, usually, found free; divided by two sinuate and perforate curved septa. Valve linear or oblong, sometimes with sinuate sides, and with a pseudoraphe and transverse punctate lines.
Chromatophores granular.
{37}GRAMMATOPHORA MARINA (LYNG.) KUETZ.
Valve linear-elliptical, with smooth apices. Septum with a wide undulation near its origin, thence straight and incrassate at the end. Striæ in quincunx, 18-21 in 10 µ.
_Diatoma marinum_ Lyngbye.
Blue clay. Along the coast.
Pl. 8, Figs. 17 and 18.
GRAMMATOPHORA MARINA VAR. SUBTILISSIMA (BAIL.) V. H.
Valve linear, slightly constricted near the smooth apices. Septum undulated near its origin and then straight, incrassate at the end. Puncta in quincunx very subtle, 34-36 in 10 µ.
_Grammatophora subtilissima_ Bail.
Grammatophora oceanica var. subtilissima (Bail.) V. H., according to De Toni. G. marina and G. oceanica are united by some authors; the latter has more subtle striæ.
Along the coast.
Pl. 8, Figs. 13 and 14.
GRAMMATOPHORA SERPENTINA RALFS
Valve linear-elliptical, long, measuring to 150 µ (De Toni); smooth at the apices. Septum with numerous undulations and hooked at the apex. Puncta in quincunx, 17 in 10 µ.
Along the coast.
Pl. 8, Fig. 21.
GRAMMATOPHORA ANGULOSA VAR. HAMULIFERA (KUETZ.) GRUN.
Frustule nearly quadrate; valve with rounded but not smooth apices. Septum bent into a sharp angle near its origin and ending in a broad hook. Puncta in transverse rows, 14 in 10 µ.
Along the coast.
Pl. 8, Figs. 15 and 16.
GRAMMATOPHORA ISLANDICA EHR.
Frustule oblong; valve elliptical-lanceolate. Septum robust with several undulations and hooked at the end. Pseudoraphe distinct; transverse rows of puncta, 10 in 10 µ.
Reported by Kuetzing in the Atlantic Ocean and by Kain at Belmar, N. J. I have not found it on our coast and I believe, in some cases, it has been confused with _G. angulosa_ var. _hamulifera_. The figure is drawn from an Iceland form in H. L. Smith T. S., 186.
Pl. 8, Figs. 19 and 20.
STRIATELLA AG. (1832)
(dim. of stria, referring to the lines on the frustule)
Frustules tabulate, adnate in short, stipitate filaments, scarcely siliceous, divided into partitions, septate or partly so at alternate ends.
{38}STRIATELLA UNIPUNCTATA (LYNG.) AG.
Frustules with numerous bent septa extending the entire length. Valve lanceolate, somewhat unsymmetrical, subtly punctate, with pseudoraphe quite distinct.
"The specific name is derived from the appearance of the endochrome which in the living specimen is invariably collected in a central mass with slender threads radiating in all directions toward the cell-wall" (Wm. Sm.). Pyrenoids cuneate, in the centre of the endochrome, numerous.
Long Island Sound and along the coast.
Pl. 8, Figs. 22 and 23.
STRIATELLA INTERRUPTA (EHR.) HEIB.
Frustules quadrangular, with robust alternate septa extending to the middle. Puncta in quincunx, 22 in 10 µ.
_Tessella interrupta_ Ehr.
Very rare along the coast.
Pl. 8, Fig. 24. (From a form found at Stonington, Conn.)
ATTHEYA WEST (1860)
(named after Thomas Atthey)
Frustules quadrangular, tabulate, with numerous annuli. Valve elliptical-lanceolate, with a pseudoraphe and a central punctum. Extending from each end is a strong spine half as long as the valve.
ATTHEYA DECORA WEST
The only species. Diagnosis of the genus. The valves are imperfectly siliceous, scarcely visible in balsam.
Very local. Abundant at Shark River, N. J.
Pl. 8, Fig. 25.
(_b_) MERIDIONEÆ
Valve symmetrical in zone and valve view along the sagittal line, but asymmetrical to the transverse axis, cuneate. In zone view sometimes with wedge-shaped septa. Valve finely striated, without central and usually without terminal nodules; a pseudoraphe present.
_Licmophora._--Frustules cuneate in stipitate fan-shaped fascicles.
_Meridion._--Frustules cuneate in spiral fascicles.
LICMOPHORA AG. (1827)
(licmos, a fan, and phoreo, I bear)
Frustules wedge-shaped, joined together into fan-shaped, stipitate fascicles. Valve cuneate, rounded at both ends, septate. Chromatophores granular, round or oval in our species.
{39}ANALYSIS OF SPECIES
(In accordance, so far as it relates to our species, with the classification of C. Mereschkowsky, Diagnoses of New Licmophoræ, Nuova Notarisia, 1901.)
Placatæ--valve narrow, striæ very fine, septa superficial flabellata
Dubiæ--valve bacilliform, septa shallow, frustule with
thick walls ovulum
Paradoxæ--valve with lower end produced, striæ fine, paradoxa
pseudoraphe distinct, septa deep gracilis
tincta
baileyi ?
Lyngbyeæ--valve narrow, attenuated at both ends, distinct,
septa deep lyngbyei
Peristriatæ--valve broad, pseudoraphe wide, striæ robust ehrenbergii
LICMOPHORA FLABELLATA (CARM.) AG.
Frustule elongate, narrow; valve narrow, lanceolate-cuneate, enlarged at the base; striæ very fine, 30 in 10 µ.
_Echinella flabellata_ Carm.
_Licmophora splendida_ Wm. Sm.
Common along the coast.
Pl. 9, Figs. 1 and 2.
LICMOPHORA OVULUM MER.
Valve ovate, attenuated to the rounded inferior apex; pseudoraphe indistinct, striæ fine, 24 in 10 µ. Zone view broad, cuneate, angles rounded, inferior apex broad; frustule robust, septa superficial, straight. (Mereschkowsky, in part.)
Atlantic City. Common.
Pl. 9, Figs. 8 and 9.
LICMOPHORA PARADOXA (LYNG.) AG.
Frustule broad, with rounded angles; septa curved; valve ovate, inferior apex produced. Pseudoraphe distinct; striæ varying from 25 below to 30 above in 10 µ.
_Echinella paradoxa_ Lyng.
_Rhipidophora paradoxa_ Kuetz.
Along the coast.
Pl. 9, Figs. 6 and 7.
LICMOPHORA GRACILIS (EHR.) GRUN.
Frustule cuneate, narrow, with sinuate margin; valve clavate, linear at the base; striæ, 20 to 22 in 10 µ.
New Rochelle. Along the coast.
Pl. 9, Fig. 11.
LICMOPHORA GRACILIS VAR. ELONGATA (KUETZ.) DE TONI
As in the type, but more graceful and with deeper septa.
_Rhipidophora elongata_ Kuetz.
Along the coast. Not common.
Pl. 9, Figs. 12 and 13.
{40}LICMOPHORA TINCTA (AG.) GRUN.
Frustules cuneate, narrow, usually found in twos. Valve clavate, hyaline, rather broad at the base; septa moderately deep; pseudoraphe indistinct; striæ, 27 at the base, 30 in the middle and 33 at the apex in 10 µ.
_Gomphonema tinctum_ Ag.
Along the coast. Abundant from about the middle of July to the middle of August.
Pl. 9, Figs. 14 and 15.
LICMOPHORA BAILEYI (EDW.) GRUN.
Frustule broadly cuneate or with convex margins, rarely almost orbicular; valve spatulate or ovate with slender, produced base; septa very deep; pseudoraphe distinct; striæ 20 in 10 µ.
_Podosphenia baileyi_ (Edw.) Lewis.
Long Island Sound and upper coast of New Jersey.
This form is placed in a doubtful position by Mereschkowsky. As it corresponds more closely to the Paradoxæ, it is placed here provisionally. The girdle face and apex of the valve are round, the pseudoraphe is distinct and the septa deep, but the stipe is short.
Pl. 9, Fig. 10 and Pl. 38, Figs. 3 and 4.
LICMOPHORA LYNGBYEI (KUETZ.) GRUN.
Frustule cuneate, slightly rounded at the angles. Valve oblanceolate; pseudoraphe distinct; septa deep; striæ, 12 in 10 µ below, and 16 in 10 µ above.
_Podosphenia lyngbyei_ Kuetz.
Along the coast.
Pl. 9, Figs. 3 and 4.
LICMOPHORA EHRENBERGII (KUETZ.) GRUN.
Frustule cuneate, broad. Valve obovate-lanceolate; pseudoraphe wide; striæ coarse, 8 in 10 µ, moniliform.
_Podosphenia ehrenbergii_ Kuetz.
Along the coast.
Pl. 9, Fig. 5.
MERIDION AG. (1824)
(merizo, I divide)
Frustules in zone view cuneate, adnate in circular or spiral fasciæ, at length becoming free. Valve symmetrical with respect to the longitudinal axis, more or less cuneate; costæ and striæ transverse.
Chromatophores numerous, small, elongated, in irregular rows on the zone (Pfitzer).
MERIDION CIRCULARE (GREV.) AG.
Transverse costæ coarse, variable in number and distance apart, sometimes interrupted or indistinct; striæ interstitial, 16 in 10 µ.
In springs and small streams of pure water.
_Echinella circularis_ Grev.
{41}Meridion constrictum Ralfs, sometimes given as a variety of M. circulare, differs only in the constriction below the apex. The two kinds of frustules are usually found growing together and as the variation is often extremely slight they are here included under the earlier name.
Pl. 10, Figs. 1, 2 and 3.
Fig. 1 represents the constricted form which is the more common. Fig. 3 is a sporangial form.
The sporangial frustules vary in shape and size, some being long and slender, others clavate, but they are all more or less tumid in the middle, with costæ more indefinite than in perfect valves. All gradations occur, one end becoming shorter until the valve has the shape of the variety known as constrictum. It would seem, therefore, that the non-constricted form is a passage from the sporangial to the smaller or adult form, or is of no specific importance. All forms are found living together. The adult frustules are the smaller ones; it is from them that the sporangia are produced.
Meridion intermedium H. L. Smith (Amer. Quart. Mic. Jour., Vol. 1, p. 12) is characterized by less evident costæ and is more delicate in general appearance. Some forms are capitate and others are not. Prof. Smith compares the M. intermedium with Peronia erinacea Bréb. and Arnott which he has named M. erinaceum, hitherto found only in Europe, and points out the relation of the two forms to Licmophora. An examination of the H. L. S. type slides of the two diatoms proves that Peronia has very delicate costæ and a distinct pseudoraphe not noticeable in Meridion. On the slide of Peronia are frustules exactly similar to certain of the sporangial variations of M. circulare.
The fan-like arrangement of Licmophora, the marine form, and the circular chains of Meridion, the fresh-water genus, are similar. Both are stipitate at the beginning of their growth.
(_c_) FRAGILARIEÆ
Divided into three sections:
_Diatominæ._--Valve circular, elliptical to linear, quadrate or cruciform, with transverse costæ; without raphe, a pseudoraphe sometimes wanting.
_Fragilariinæ._--Valve elongate, with small central and terminal elevations, without costæ but with transverse punctate striæ; without genuine central nodule.
_Eunotiinæ._--Valve lunate; a raphe sometimes partially formed with terminal nodules near the edges.
DIATOMINÆ
_Diatoma._--Frustules in filaments. Valve linear or elliptical, costate.
_Plagiogramma._--Frustules in fasciæ or free. Valve costate.
_Opephora._--Valve costate, with an inner punctate stratum.
DIATOMA DE CANDOLLE (1805) em. HEIB. (1863)
(diatemno, I cut in two)
Frustules oblong or quadrate, adnate in filaments, attached by alternate angles and finally separating. Valve linear or elliptical, with transverse costæ and rows of puncta and a pseudoraphe.
Chromatophores large granules without definite arrangement. (See Pl. 40, Fig. 11.)
{42}DIATOMA VULGARE BORY.
Valve elliptical-lanceolate, with apices sometimes rostrate or capitate; pseudoraphe narrow; costæ, 5 in 10 µ.
Common everywhere in pure fresh water and extremely variable.
Pl. 10, Figs. 9 and 10.
Var. elongatum (Ag.) = var. ehrenbergii (Kuetz.)--elliptical-lanceolate, constricted near the apex.
Var. grande (Wm. Sm.) Grun.--linear, elongated, constricted near the apices.
Pl. 10, Fig. 4.
Both of these varieties, with numerous intermediate forms, are abundant near Newtown Square. Varieties of Grunow, known as breve, ovate-lanceolate; productum, ovate-lanceolate with produced apices; capitulatum, lanceolate with capitate extremities, are mingled together in the same gathering.
DIATOMA ANCEPS (EHR.) KIRCHN.
Valve linear with rostrate apices; costæ robust; striæ delicate, 20 in 10 µ. Zone view quadrangular.
Common in fresh water.
Pl. 10, Figs. 5 and 6. Fig. 11, Pl. 40, shows frustules containing the nuclei and chromatophores.
DIATOMA HIEMALE (LYNG.) HEIB.
Valve ovate-lanceolate to lanceolate; apices obtuse, not produced. Costæ not numerous, robust; striæ moniliform. Zone view quadrate, the costæ as septa deeply dividing the valve into convex elevations.
Common in springs.
Pl. 10, Figs. 7 and 8.
In all species of Diatoma a punctum, or pore, is observed, usually at alternate ends of the two valves, by means of which a communication exists between adjoining frustules and causes them to adhere in zig-zag chains when partially separated.
PLAGIOGRAMMA GREV. (1859)
(plagios, on the side, and gramma, a letter)
Frustules quadrangular, adnate in fasciæ, or free. Valve linear, elliptical, or elliptical-lanceolate, divided by two or more median and two terminal costæ or with a central and two terminal hyaline spaces.
Valve with two median and two terminal costæ:
Linear, pseudoraphe distinct pygmæum
Linear, with striæ at the ends wallichianum
Ovate-lanceolate obesum
Valve without costæ but with central and terminal
nodules: pseudoraphe absent tessellatum
{43}PLAGIOGRAMMA PYGMÆUM GREV.
Valve linear-elliptical; pseudoraphe distinct; rows of granules transverse, usually six in each compartment, moniliform, three on each side.
Blue clay. Not common.
Pl. 10, Fig. 13.
PLAGIOGRAMMA WALLICHIANUM GREV.
Valve linear, rounded at the ends; pseudoraphe absent; transverse rows of granules, six or seven in each compartment, and two or three rows of smaller granules at each end.
Blue clay. Not common.
Pl. 10, Fig. 14.
PLAGIOGRAMMA OBESUM GREV.
Valve rhombic-lanceolate, the costæ scarcely visible; pseudoraphe rather wide; rows of granules, about seven in each compartment, slightly radiating.
Blue clay. Not common.
Pl. 10, Fig. 12.
PLAGIOGRAMMA TESSELLATUM GREV.
Valve elliptical-lanceolate; central space transversely elliptical to the major axis, half the diameter of the valve; terminal spaces more or less circular or ovate. Granular markings large, quadrangular, in transverse rows. Pseudoraphe not distinct. As the central space does not reach the margin, it is a question whether this form is a Plagiogramma or a new genus.
Blue clay. Rare.
Pl. 10, Fig. 11.
OPEPHORA PETIT (1888)
(ope, an opening, and phoreo)
Frustule rectangular. Valve cuneiform, linear or elliptical-lanceolate, with broad, transverse striæ and a well-defined pseudoraphe or median area.
The genus "portant des stries en forme de boutonnières," as Petit remarks, is quite near Fragilaria, under which the species here described were originally included. (See Schmidt's Atlas, Pl. 298, where numerous forms of F. pinnata are figured.)
OPEPHORA SCHWARTZII (GRUN.) PETIT
Valve obovate-lanceolate or nearly linear with rounded apices; striæ transverse, broad, 3 or 4 in 10 µ; median area lanceolate.
An inner stratum, with puncta in transverse rows, is apparent.
Blue clay. Not uncommon. Variable in size.
Pl. 10, Figs. 16 and 19.
OPEPHORA PACIFICA (GRUN.) PETIT
Valve linear, oblong, with rounded apices. Median area linear, narrow; striæ punctate.
Blue clay.
Pl. 10, Fig. 18.
Petit (Diat. Cap Horn) in his diagnosis states that the valves are cuneiform, but they are not always so.
{44}OPEPHORA PINNATA VAR. LANCEOLATA N. VAR.
Valve lanceolate; costæ slightly radiate, punctate; median area broad, lanceolate.
Differs from O. pinnata in outline, radiation of the costæ and median area.
Blue clay. Rare.
Pl. 10, Fig. 17.
FRAGILARIINÆ
_Fragilaria._--Frustules in fasciæ. Valve with transverse striæ. Pseudoraphe indistinct.
_Rhaphoneis._--Striæ radiate; pseudoraphe distinct.
_Dimerogramma._--Pseudoraphe broad.
_Trachysphenia._--Valve cuneiform.
_Synedra._--Valve elongate.
_Asterionella._--Frustules in star-shaped clusters.
FRAGILARIA (LYNG.) RAB.
(fragilis, because of the fasciæ easily breaking up)
Frustules rectangular, adnate in fasciæ, soon breaking up. Valve lanceolate, oblong or elliptical in general outline, with convex or sinuate margins; without costæ; pseudoraphe narrow or indistinct; striæ transverse. Chromatophores vary according to species. In some they consist of four bands on the valves; in others they are granular (Mereschkowsky).
Brun divides the genus into two sections, Fragilaria proper and Staurosira. The former, with an indistinct pseudoraphe, includes the species virescens, arctica, undata and linearis, while the latter, with distinct pseudoraphe, includes capucina, harrisonii, construens and parasitica.
FRAGILARIA VIRESCENS RALFS
Frustules in long fasciæ. Valve elliptical-lanceolate, obtuse at the apices; pseudoraphe indistinct; striæ, 17 in 10 µ, punctate.
Very common in springs and pure streams. The fasciæ are often a foot or more in length.
Pl. 10, Figs. 20 and 21.
FRAGILARIA ARCTICA GRUN.
Valve oblong or elliptical, 10 µ in length; striæ subtle, with coarse, short striæ at intervals on the margin and evident in zone view.
Marine. Common at Cape May, N. J.
Pl. 10, Figs. 22 and 23.
FRAGILARIA UNDATA WM. SM.
Valve in general outline linear-elliptical, with extremities produced; striæ subtle; pseudoraphe distinct.
Fresh water.
Pl. 10, Figs. 24, 25, 27, 28 and 29.
{45}FRAGILARIA LINEARIS CSTR.
Valve linear, with rounded apices; striæ subtle; pseudoraphe indistinct.
Marine. Cape May.
Pl. 10, Fig. 37. Fig. 36 is an indeterminate form occasionally found in the blue clay.
FRAGILARIA CAPUCINA VAR. MESOLEPTA RAB.
Valve linear, constricted at the hyaline middle; apices slightly produced; striæ, 17 in 10 µ. Quite variable in size.
Schuylkill River. Morrisville (Keeley).
Pl. 10, Fig. 34.
FRAGILARIA HARRISONII (WM. SM.) GRUN.
Frustules rectangular, solitary or in twos. Valve cruciform; pseudoraphe narrow, lanceolate; striæ robust, radiating in the middle, composed of confluent puncta, larger at the circumference.
Blue clay.
Pl. 10, Fig. 31.
FRAGILARIA CONSTRUENS (EHR.) GRUN.
Valve in general outline lanceolate, with produced apices; pseudoraphe lanceolate, distinct or broad; striæ subtle, 15 in 10 µ. L. of valve, 10-45 µ.
_Staurosira construens_ Ehr.
_Odontidium tabellaria_ Wm. Sm.
Blue clay.
Pl. 10, Fig. 30.
FRAGILARIA PARASITICA (WM. SM.)
Frustules solitary or in twos. Valve lanceolate, sometimes constricted in the middle; pseudoraphe wide, lanceolate; striæ subtle. Parasitic on other diatoms.
_Odontidium parasiticum_ Wm. Sm.
Not common. Media (Palmer).
In the constricted form it is known as F. construens var. binodis (Ehr.) Grun.
Pl. 10, Fig. 35.
An examination of the synonymy of the species of Fragilaria will convince the student of the difficulty of determining the correct name even in well-known forms. If all of the species of Fragilaria proper have granular chromatophores, and all of Staurosira are placcochromatic, a satisfactory division can be made, but so long as these facts are not known in all species, and as authors have repeatedly confused the two divisions, the nomenclature will be uncertain. F. harrisonii is probably in any case to be separated from the others. De Toni includes it under its original name of Odontidium, which genus he places near to Diatoma. The number of species in our locality is too limited to render further discussion of any value.
{46}RHAPHONEIS EHR. (1844)
(rhaphis, a needle)
Frustule in zone view linear. Valve lanceolate or elliptical-lanceolate; pseudoraphe distinct; striæ radiating, moniliform.
RHAPHONEIS AMPHICEROS EHR.
Valve lanceolate, broad, with apices produced; striæ in curved lines, moniliform, the large granules in longitudinal lines.
Blue clay.
Pl. 10, Fig. 38.
RHAPHONEIS AMPHICEROS VAR. RHOMBICA GRUN.
Valve as in type form but shorter, with larger and more remote granules.
Blue clay.
Pl. 10, Figs. 39 and 40.
RHAPHONEIS BELGICA VAR. INTERMEDIA GRUN.
Valve lanceolate, rostrate; granules in longitudinal and nearly transverse, not radiating, lines.
Absecon, N. J.
Pl. 10, Fig. 41.
DIMEROGRAMMA RALFS (1861)
(dis, two, meros, a part, gramma, a letter)
Frustules quadrangular, inflated at the angles, in fasciæ. Valve ovate or lanceolate; striæ moniliform, transverse or slightly radiate; median area or pseudoraphe broad, lanceolate.
DIMEROGRAMMA MARINUM (GREG.) RALFS
Valve lanceolate or linear and inflated in the middle; striæ moniliform, transverse or slightly radiate; median area linear or lanceolate, sometimes not reaching the smooth extremities; striæ, 8 in 10 µ.
Pl. 12, Figs. 9 and 10.
Fig. 9 differs in its lanceolate outline, in having four puncta on each side in a row, and in the striæ which are radiate.
DIMEROGRAMMA SURIRELLA (EHR.) GRUN.
Valve elliptical-lanceolate, with rounded apices; striæ moniliform, radiate; pseudoraphe narrow, lanceolate.
Blue clay.
Pl. 12, Fig. 11.
{47}DIMEROGRAMMA MINUS (GREG.) RALFS
Valve rhombic-lanceolate; striæ punctate, radiate; pseudoraphe lanceolate; apices smooth.
Blue clay. Along the coast.
Pl. 12, Figs. 12, 13, 14.
TRACHYSPHENIA PETIT (1877)
(trachys, rough, and sphen, a wedge)
Frustules rectangular. Valve cuneiform with coarse puncta in transverse and longitudinal lines; pseudoraphe narrow, linear. One species only.
TRACHYSPHENIA AUSTRALIS PETIT
Characters of the genus. Valve small; puncta, 6 in 10 µ. Allied to Dimerogramma.
Shark River, N. J. Rare.
Pl. 12, Fig. 15.
SYNEDRA EHR. (1830)
(synedrion, a sitting together)
Frustules adnate in small stipitate clusters or free. Valve elongate, linear or linear-lanceolate; pseudoraphe distinct; costæ absent.
The genus Synedra has few distinctive characters. As Brun remarks (Diat. des Alpes et du Jura, p. 122), the dilatation of the extremities and the pseudo-nodule are of little value in classification, as the intermediate forms are so numerous. Fragilaria occurs in very long ribbons or fasciæ, Synedra in short fasciæ or radiating clusters. Fragilaria is seldom longer than three or four times the width, while Synedra is nearly always so. The former has fine, often subtle, markings and narrow pseudoraphe, while the latter has coarser punctate striæ and a more distinct pseudoraphe.
Chromatophores usually consist of two bands, one on each of the valves. Karsten states that in the marine forms the chromatophores are oval or polygonal discs, each of which usually encloses a pyrenoid.
SYNEDRA ULNA (NITZSCH) EHR.
Frustules solitary or in twos. Valve 150-250 µ in length, linear or linear-lanceolate, with rostrate apices; striæ, 9 in 10 µ.
Common in rivers and streams.
Pl. 11, Figs. 4, 7 and 11 (?).
Frequently interrupted in the middle. The distinction made by Wm. Smith as to the presence or absence of the central blank space is probably not necessary, as both forms are found which are otherwise identical.
Fig. 5 represents the formation of a sporangial frustule which differs from the usual form in its inflated ends prolonged into rostrate apices. Figs. 1 and 6 are sporangial frustules.
{48}SYNEDRA BICEPS (KUETZ.) SCHMIDT
Valve sublanceolate, inflated at the ends, apices rounded; central space not always distinct; pseudoraphe narrow; striæ radiate at the ends.
This is not Kuetzing's species, if the descriptions and figures are accepted, nor is it H. L. Smith's Type No. 545, which is S. ulna var. danica, nor is it S. biceps Wm. Smith, but it is exactly Schmidt's form (Atlas, Pl. 303, Figs. 10-15).
Schuylkill River.
Pl. 11, Fig. 3.
SYNEDRA DANICA KUETZ.
Valve lanceolate, suddenly constricted at the rounded apices; central space frequently absent.
Very common in streams.
Pl. 11, Fig. 2.
The figure represents an unusually large form. It differs from S. ulna only in its apices.
SYNEDRA CAPITATA EHR.
Valve long, linear, dilated into triangular acute apices; pseudoraphe distinct; striæ radiate at the ends.
Blue clay.
Pl. 11, Fig. 8.
SYNEDRA ACUS KUETZ.
Valve very narrow, lanceolate, acicular, with obtuse apices.
Common in the Schuylkill River.
Pl. 11, Figs. 9 and 18.
SYNEDRA GOULARDI BRÉB.
Valve constricted in the middle; apices sub-acute, sometimes slightly rostrate or capitate; central space evident.
Neshaminy Creek (Palmer). Blue clay. Crum Creek.
Pl. 11, Figs. 12 and 13.
SYNEDRA PULCHELLA (RALFS) KUETZ.
Valve lanceolate, tapering to the sub-acute, rostrate or slightly capitate apices; dilated at the central hyaline space; pseudoraphe distinct. Very variable in size.
Crum Creek. Schuylkill River. Rather common.
Pl. 11, Figs. 14, 15, 16.
SYNEDRA PULCHELLA VAR. ABNORMIS MACCHIATI?
Valve as in type form, except that one end is curved like a beak, as in S. hamata Wm. Sm., which it resembles.
Not uncommon in the Schuylkill River.
Pl. 11, Fig. 17.
SYNEDRA OXYRHYNCHUS VAR. UNDULATA GRUN.
Valve linear-lanceolate with produced rostrate apices, asymmetrical, sigmoid; pseudoraphe narrow; pseudo-nodule large.
Common in fresh water.
Pl. 12, Fig. 1.
{49}SYNEDRA PULCHELLA VAR. FLEXELLA N. VAR.
Frustule slightly attenuated at the ends, truncate, somewhat tumid in the middle and flexed. Valve lanceolate, with obtuse or subcapitate apices and with two almost imperceptible constrictions at the middle producing a tumid appearance; pseudoraphe distinct; pseudo-nodule absent. L. 56 µ; striæ, 14-16 in 10 µ.
Some valves are bent and incised on one side. The outline of the valve is that of pulchella.
Common at Newtown Square.
Pl. 12, Fig. 2.
SYNEDRA RADIANS KUETZ.
Frustules linear, in small fasciæ. Valve 34 µ in length, linear, with apices rostrate, obtuse, sometimes slightly capitate; pseudoraphe distinct; striæ about 20 in 10 µ.
Fresh water.
Pl. 10, Figs. 32 and 33.
There is difficulty in recognizing S. radians K. as described and figured by different authors. On Plate 12, Fig. 8, I have drawn a specimen from H. L. Smith's Type Slide No. 574, labelled S. radians Kuetz., not Wm. Smith, which, however, corresponds closely to Smith's figure (Brit. Diat. 1, Pl. 11, Fig. 89). De Toni gives S. radians Kuetz. as equivalent to S. tenera Wm. Sm. Van Heurck's figure of S. radians, and also the figure of ulna var., said to be synonymous with H. L. Smith's S. radians, which does not correspond to the specimens on Smith's slide in my possession, are confusing. In Van Heurck's Synopsis the striæ are said to be 16 or 17, while De Toni describes them as subtle and from 17 to 24 in 10 µ. The length is quite variable.
Several species of Synedra resemble S. radians in the mode of growth, as they are adnate at first, in short bands, the frustules being sessile on other plants or objects, attached at the terminal nodules which, although scarcely visible in most forms, are probably present in all. The frustules are not closely connected at the free end, and soon become entirely detached.
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The Diatomaceæ of Philadelphia and VicinityChapter II: Preface (2)
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