Chapter XI: Literature and History (29)

Cortical shell thorny, with eight long radial spines, lying in two crossed diagonal planes, opposite in pairs. These eight horns are the prolongations of the anterior and posterior edges of each wing, on both sides of its lateral plane; they are of great importance, as appearing (by heredity) in many other Pylonida (_e.g._, in _Tetrapyle octacantha_). Lateral crest of each wing without spines, slightly convex.

_Dimensions._--Length of the medullary shell 0.07, breadth 0.05; length of each lateral wing 0.15, breadth 0.06.

_Habitat._--Pacific, central area, Stations 270 to 274, surface.

10. _Amphipyle cladoceros_, n. sp.

Cortical shell spiny, with eight strong ramified radial spines, lying in the same two crossed diagonal planes as in the preceding species. It differs from this in the ramification of the eight horns, which bear six to nine irregular lateral branches, partly simple, partly bifurcate. Lateral crest of each wing spiny, convex.

_Dimensions._--Length of the medullary shell 0.06, breadth 0.045; length of each lateral wing (without spines) 0.2, breadth 0.09.

_Habitat._--Indian Ocean, surface, Madagascar, Rabbe.

11. _Amphipyle decaceros_, n. sp.

Cortical shell spiny, with ten strong conical spines; eight lying in two crossed diagonal planes, in the same disposition (opposite in pairs) as in both preceding species; two others opposite on the poles of the principal axis, arising from both poles of the medullary shell (columella-beams). Spiny crest of the wings convex, semilunar.

_Dimensions._--Length of the medullary shell 0.04, breadth 0.03; length of each lateral wing 0.12, breadth 0.04.

_Habitat._--North Atlantic, Station 354, surface.

12. _Amphipyle dodecaceros_, n. sp.

Cortical shell thorny, with twelve large cylindrical spines; eight lying in two crossed diagonal planes, in the same disposition (opposite in pairs) as in the three preceding species; four others crossed at right angles in the lateral plane, two opposite on the poles of the principal, two on the poles of the lateral axis (the latter shorter). Both lateral wings nearly rhombic.

_Dimensions._--Length of the medullary shell 0.06, breadth 0.04; length of the lateral wings 0.15, breadth 0.05.

_Habitat._--South Atlantic, coast of Brazil, Rabbe, surface.

{644}13. _Amphipyle callizona_, n. sp. (Pl. 9, fig. 6).

Cortical shell smooth, with sixteen strong and short conical spines, lying opposite in pairs in two crossed diagonal planes; each lateral wing four-sided prismatic, its lateral face concave, both ends truncated, and each end provided with four divergent spines. Length of the wings somewhat greater, but breadth smaller, than that of the medullary shell.

_Dimensions._--Length of the medullary shell 0.07, breadth 0.05; length of the lateral wings 0.09, breadth 0.03.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

Genus 283. _Tetrapyle_,[322] J. Müller, 1858, Monatsber. d. k. preuss. Akad. d. Wiss. Berlin, p. 154.

_Definition._--#Pylonida# with trizonal lentelliptical medullary shell, surrounded by two crossed latticed cortical girdles, one smaller (primary) transverse, and one larger (secondary) lateral girdle. Four gates between the two cortical girdles simple, without a sagittal septum.

The genus _Tetrapyle_, till 1881 the only known genus of the whole family, was founded by Johannes Müller in 1858, and clearly illustrated by the Mediterranean (and common cosmopolitan) _Tetrapyle octacantha_, hitherto the best known type of this family. Afterwards (1860, Monatsber. d. k. preuss. Akad. d. Wiss. Berlin, p. 832) Ehrenberg founded the genus _Schizomma_ for a nearly allied form, which exhibits only slight specific differences (compare my Monograph, 1862, p. 434). Some good remarks on the structure of this typical genus and its relations to other Pylonida are to be found in Richard Hertwig's Organismus, &c., 1879, p. 52, but the true trizonal structure of the medullary shell in this genus was not recognised by him, so that his description agrees more with _Dizonium_. We confine here the genus _Tetrapyle_ to those Pylonida for which _Tetrapyle octacantha_ of J. Müller remains the determining type; the cortical shell is composed only of two perfect lattice-girdles (the transverse and lateral), between which four wide gates remain open. This structure is similar to that of _Dizonium_; but whilst here the medullary shell is a simple central chamber, in _Tetrapyle_ it is a complete trizonal or _Larnacilla_-shell.

Subgenus 1. _Tetrapylissa_, Haeckel.

_Definition._--Cortical shell smooth or thorny, but without large, symmetrically disposed spines.

{645}1. _Tetrapyle circularis_, n. sp. (Pl. 9, fig. 8).

Cortical shell rough, without radial spines. Lateral girdle circular, therefore the longitudinal axis is equal to the transverse. Four gates kidney-shaped, twice as broad as high. In each half wing of the transverse girdle six to seven longitudinal rows of irregular, roundish pores.

_Dimensions._--Medullary shell 0.05 long, 0.04 broad; cortical shell 0.15 long and broad; gates 0.04 high, 0.08 broad.

_Habitat._--Pacific, central area, Station 263, surface.

2. _Tetrapyle circopyle_, n. sp.

Cortical shell smooth, without radial spines. Lateral girdle elliptical, one and a half times as long as broad. Four gates nearly circular. On each half wing of the transverse girdle four to five rows of subregular, circular pores. (Similar to _Tetrapyle pleuracantha_, Pl. 9, fig. 9, but without lateral spines.)

_Dimensions._--Medullary shell 0.04 long, 0.03 broad; cortical shell 0.16 long, 0.11 broad; gates 0.05 high and broad.

_Habitat._--Pacific, central area, Station 268, depth 2900 fathoms.

3. _Tetrapyle quadriloba_, Haeckel.

_Tetrapyle quadriloba_, Haeckel, 1862, Monogr. d. Radiol., p. 436.

_Schizomma quadrilobum_, Ehrenberg, 1860, Monatsber. d. k. preuss. Akad.
d. Wiss. Berlin, p. 815; Abhandl. d. k. Akad. d. Wiss. Berlin, 1872, Taf.
x. figs. 12-14.

Cortical shell thorny, without regular, radial spines. Lateral girdle elliptical, one and a half times as long as broad. Four gates nearly circular. On each half wing of the transverse girdle three to four rows of irregular, roundish pores.

_Dimensions._--Medullary shell 0.06 long, 0.04 broad; cortical shell 0.16 long, 0.1 broad; diameter of the gates 0.05.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Indian Ocean, Pacific, surface.

4. _Tetrapyle nephropyle_, n. sp.

Cortical shell thorny, without radial spines. Lateral girdle elliptical, one and a third times as long as broad. Four gates kidney-shaped, nearly twice as broad as high, with two prominent polar spines on the poles of the longitudinal axis of the medullary shell. On each half wing of the transverse girdle three to four longitudinal rows of large roundish pores.

_Dimensions._--Medullary shell 0.06 long, 0.04 broad; cortical shell 0.24 long, 0.18 broad; gates 0.05 high, 0.08 broad.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{646}5. _Tetrapyle cardiopyle_, n. sp.

Cortical shell rough, without radial spines. Lateral girdle with hexagonal contour, nearly cylindrical in the middle part, conical at both ends, twice as long as broad. Four gates heart-shaped, about as high as broad. On each half wing of the transverse girdle three to four longitudinal rows of subregular, polygonal pores.

_Dimensions._--Medullary shell 0.04 long, 0.02 broad; cortical shell 0.14 long, 0.07 broad; gates 0.05 high and broad.

_Habitat._--Eastern Tropical Atlantic, Station 348, depth (2450) fathoms.

Subgenus 2. _Tetrapylura_, Haeckel.

_Definition._--Cortical shell armed with large, symmetrically disposed spines.

6. _Tetrapyle fusiformis_, n. sp.

Cortical shell rough, with two strong conical spines on the poles of the longitudinal axis. Lateral girdle spindle-shaped, one and a half times as long as broad. Four gates kidney-shaped, twice as broad as high. On each half wing of the transverse girdle four to five longitudinal rows of irregular, roundish pores.

_Dimensions._--Medullary shell 0.07 long, 0.05 broad; cortical shell 0.18 long, 0.12 broad; gates 0.04 high, 0.08 broad.

_Habitat._--Western Tropical Atlantic, Mexican Gulf Stream, depth 1500 fathoms, Schaffner.

7. _Tetrapyle pleuracantha_, n. sp. (Pl. 9, fig. 9).

Cortical shell rough, with two strong conical spines on the poles of the transverse axis. Lateral girdle elliptical, one and half times as long as broad. Four gates subcircular or nearly hexagonal, about as high as broad. On each half wing of the transverse girdle three to four longitudinal rows of very irregular pores.

_Dimensions._--Medullary shell 0.045 long, 0.035 broad; cortical shell 0.16 long, 0.11 broad; gates 0.045 high and broad.

_Habitat._--Western Tropical Pacific, off New Guinea, Stations 222 to 224, depths 1850 to 2450 fathoms.

8. _Tetrapyle transversa_, n. sp.

Cortical shell thorny, with two strong conical lateral spines on the poles of the transverse axis. Lateral girdle elliptical, only four-fifths as long as broad. Four gates elliptical, twice as broad as high, with two prominent spines on the poles of the longitudinal axis of the medullary shell. On each half wing of the transverse girdle five to six longitudinal rows of subregular, roundish pores. (Similar to _Octopyle transversa_, but without sagittal septum and with stronger lateral spines.)

{647}_Dimensions._--Medullary shell 0.05 long, 0.035 broad; cortical shell 0.12 long, 0.15 broad; gates 0.03 high, 0.06 broad.

_Habitat._--Southern Pacific, Station 290, surface.

9. _Tetrapyle cruciata_, n. sp.

Cortical shell smooth, with four strong radial spines, two on the poles of the longitudinal axis, two on the poles of the transverse axis. Lateral girdle nearly circular, quite as long as broad. Four gates kidney-shaped, twice as broad as high. On each half wing of the transverse girdle five to six rows of subregular, circular pores.

_Dimensions._--Medullary shell 0.06 long, 0.05 broad; cortical shell 0.16 long and broad; gates 0.05 high, 0.09 broad.

_Habitat._--South Atlantic, Station 330, surface.

10. _Tetrapyle staurophora_, n. sp.

Cortical shell thorny, with four strong radial spines, two on the poles of the longitudinal axis, two on the poles of the transverse axis. Lateral girdle elliptical, one and a half times as long as broad. Four gates nearly hexagonal, quite as high as broad. On each half wing of the transverse girdle three to four longitudinal rows of irregular, roundish pores.

_Dimensions._--Medullary shell 0.06 long, 0.04 broad; cortical shell 0.18 long, 0.12 broad; gates 0.08 high and broad.

_Habitat._--Western Tropical Atlantic, coast of Brazil, Rabbe, surface.

11. _Tetrapyle quadricornis_, n. sp.

Cortical shell thorny, with four strong horn-like curved spines, lying in the lateral plane and converging in pairs towards the poles of the transverse axis. Lateral girdle elliptical, one and a third times as long as broad. Four gates kidney-shaped, twice as broad as high. On each half wing of the transverse girdle four to five rows of irregular, roundish pores. (May be only the young form of _Pylonium quadricorne_, Pl. 9, fig. 14.)

_Dimensions._--Medullary shell 0.05 long, 0.04 broad; cortical shell 0.16 long, 0.12 broad; gates 0.03 high, 0.07 broad.

_Habitat._--Pacific, central area, Stations 270 to 274, surface.

12. _Tetrapyle tetracantha_, n. sp.

Cortical shell thorny, with four strong radial spines lying in the lateral plane and in pairs in its two crossed diagonal axes. Lateral girdle elliptical, constricted in the middle, nearly twice as long as broad. Four gates kidney-shaped, one and a half times as broad as high. On each half wing of the transverse girdle three to four longitudinal rows of large, irregular, roundish pores. {648}(Similar to _Tetrapylonium quadrangulare_, Pl. 49, fig. 15, but without a sagittal girdle and with stronger lateral spines.)

_Dimensions._--Medullary shell 0.06 long, 0.04 broad; cortical shell 0.18 long, 0.1 broad; gates 0.007 high, 0.01 broad.

_Habitat._--North Atlantic, Canary Islands, Station 353, surface.

13. _Tetrapyle quadrigata_, n. sp.

Cortical shell thorny, with four brush-like groups of radial spines on the four diagonal corners of the lateral plane, opposite in pairs in diagonal axes. Lateral girdle nearly square, somewhat broader than long. Four gates kidney-shaped, twice as broad as high. On each half wing of the transverse girdle five to six longitudinal rows of irregular, roundish pores.

_Dimensions._--Medullary shell 0.06 long, 0.04 broad; cortical shell 0.14 long, 0.15 broad; gates 0.03 high, 0.07 broad.

_Habitat._--Pacific, central area, Station 270, surface.

14. _Tetrapyle octacantha_, J. Müller.

_Tetrapyle octacantha_, J. Müller, 1858, Abhandl. d. k. Akad. d. Wiss.
Berlin, p. 33, Taf. ii. figs. 1-6.

_Tetrapyle octacantha_, Haeckel, 1862, Monogr. d. Radiol., p. 435.

_Tetrapyle octacantha_, R. Hertwig, 1879, Organismus d. Radiol., p. 52,
Taf. iv. fig. 7, Taf. vi. figs. 2, 5, 5_a._

Cortical shell thorny, with eight long and thin, cylindrical radial spines, lying in two crossed diagonal planes, opposite in pairs. These eight characteristic diagonal spines (or "angular spines") are the same as in _Amphipyle octoceros_ and many other Pylonida, and arise as prolongations of the proximal edges of the four gates (or of the lateral wings on the eight points, where they are intersected by the edges of the lateral ring). This cosmopolitan, widely distributed and very variable species was a long time the only known species of all the Pylonida, and very accurately first described (1858) by Johannes Müller, afterwards (1879) by R. Hertwig. But in the descriptions of these authors also some different species (such as the following) may be confounded with the true typical _Tetrapyle octacantha_. The four gates of this species are transverse-elliptical or almost triangular, nearly twice as broad as high.

_Dimensions._--Length of the medullary shell 0.04, breadth 0.03; length of the cortical shell 0.18, breadth 0.13; height of the gates 0.05, breadth 0.08.

_Habitat._--Cosmopolitan; very common in all warmer seas, Mediterranean, Atlantic, Indian, Pacific, surface.

15. _Tetrapyle cladacantha_, n. sp.

Cortical shell thorny, with eight strong ramified radial spines, lying opposite in pairs in two crossed diagonal planes, as in the preceding species. It differs from this by the peculiar ramification of the eight spines, which bear two to eight simple or furcate lateral branches arising under {649}right angles (already commencing in a transitional variety of _Tetrapyle octacantha_, J. Müller, _loc. cit._, Taf. ii. figs. 5, 6). Four gates transverse-elliptical, one and a half times as broad as high.

_Dimensions._--Medullary shell 0.05 long, 0.04 broad; cortical shell 0.16 long, 0.13 broad; gates 0.05 high, 0.08 broad.

_Habitat._--Mediterranean (Corfu), Haeckel, surface.

16. _Tetrapyle pluteus_, Haeckel.

_Tetrapyle octacantha_, var., J. Müller, 1858, Abhandl. d. k. Akad. d.
Wiss. Berlin, Taf. iii. figs. 7-12.

Cortical shell thorny, with eight strong radial spines, lying opposite in pairs, as in the two foregoing species. On the distal edge each of the four triangular gates is protected by a prominent roof of lattice-work (or "pluteus"), which arises from the distal part of the lateral girdle and connects the two angular diagonal spines of each gate. Therefore the shell assumes the characteristic form very well represented by J. Müller in his fig. 11 (seen from the lateral side) and fig. 7 (seen from the pole of the principal axis).

_Dimensions._--Medullary shell 0.04 long, 0.03 broad; cortical shell 0.15 long, 0.11 broad; gates 0.045 high, 0.07 broad.

_Habitat._--Mediterranean, Atlantic, Stations 348 to 353, &c., surface.

17. _Tetrapyle turrita_, n. sp. (Pl. 9, fig. 10).

Cortical shell smooth, but with ten strong conical spines, two on the poles of the principal axis, eight others symmetrically distributed on both sides of the distal edge of the four roundish gates, and directed towards the transverse axis. Lateral girdle lanceolate, twice as long as broad.

_Dimensions._--Medullary shell 0.05 long, 0.04 broad; cortical shell 0.2 long, 0.1 broad; gates 0.05 high, 0.07 broad.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

18. _Tetrapyle dodecaceros_, n. sp.

Cortical shell thorny, with twelve stronger conical spines, eight diagonal spines in two crossed planes (as in _Tetrapyle octacantha_), and four others in the lateral plane, in pairs on both sides of a deep sagittal constriction of the quadrangular lateral girdle. Four gates kidney-shaped, twice as broad as high. On each half wing of the transverse girdle five to six longitudinal rows of irregular, roundish pores.

_Dimensions._--Medullary shell 0.06 long, 0.04 broad; cortical shell 0.18 long, 0.14 broad; gates 0.04 high, 0.08 broad.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{650}Genus 284. _Octopyle_,[323] Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Pylonida# with trizonal lentelliptical medullary shell, surrounded by two crossed, latticed, cortical girdles; one smaller (primary) transverse, and one larger (secondary) lateral girdle. Four gates between the two cortical girdles divided by a sagittal septum into eight gates.

The genus _Octopyle_ comprises those Pylonida which are distinguished from the nearly allied _Tetrapyle_ by the development of a sagittal septum, dividing the four gates of the latter into eight separate gates. The septum begins with the formation of two axial rods or columellæ, which afterwards become branched; the branches communicating one with another and with the middle parts of the lateral girdle (on the poles of the principal axis), there is formed a latticed septum in the sagittal plane, which separates more or less incompletely the right and left halves of the shell. The four gates of _Tetrapyle_ become halved by this septum, and their number doubled.

Subgenus 1. _Octopylissa_, Haeckel.

_Definition._--Cortical shell smooth or thorny, but without larger symmetrically disposed spines.

1. _Octopyle ovulina_, n. sp.

Cortical shell lentelliptical, smooth, without thorns. Lateral girdle elliptical, one and a third times as long as broad. Transverse girdle narrow, with two pores on the isthmus (or on the narrowest part of each quadrant). Sagittal septum as long as the elliptical medullary shell. Eight gates egg-shaped.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.035; length of the cortical shell 0.16, breadth 0.12.

_Habitat._--Pacific, central area, Station 274, surface.

2. _Octopyle quadrata_, n. sp.

Cortical shell thorny, nearly square, of equal length and breadth. Lateral girdle quadrangular, with rounded edges, of equal length and breadth. Transverse girdle narrow, with three pores on the isthmus. Sagittal septum of the same length as the subspherical medullary shell. Eight gates nearly circular.

_Dimensions._--Diameter of the medullary shell 0.04, of the cortical shell 0.14.

_Habitat._--Northern Pacific, Station 244, depth 2900 fathoms.

{651}3. _Octopyle subglobosa_, n. sp.

Cortical shell nearly spherical, thorny, of equal length and breadth. Lateral girdle broad, nearly circular. Transverse girdle broad, with four pores on the isthmus. Sagittal septum shorter than the subspherical medullary shell. Eight gates egg-shaped.

_Dimensions._--Diameter of the medullary shell 0.05, of the cortical shell 0.13.

_Habitat._--Southern Pacific, Station 300, depth 1375 fathoms.

4. _Octopyle transversaria_, n. sp.

Cortical shell thorny, transverse-elliptical. Lateral girdle very broad, one and a third times as broad as long. Transverse girdle also very broad, with five pores on the isthmus. Sagittal septum shorter than the elliptical medullary shell. Eight gates small, roundish.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.04; length of the cortical shell 0.11, breadth 0.15.

_Habitat._--Western Tropical Pacific, Station 224, depth 1850 fathoms.

Subgenus 2. _Octopylura_, Haeckel.

_Definition._--Cortical shell armed with large, symmetrically disposed spines.

5. _Octopyle amphistyle_, n. sp.

Cortical shell smooth, nearly spindle-shaped, about twice as long as broad. Transverse girdle broad, with four large pores on the isthmus. Lateral girdle lanceolate. Sagittal septa about as long as the medullary shell, prolonged at both poles of the principal axis into two opposite, strong, angular spines. Eight gates obliquely quadrangular.

_Dimensions._--Length of the medullary shell 0.07, breadth 0.05; length of the cortical shell 0.2, breadth 0.12.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

6. _Octopyle staurostyle_, n. sp.

Cortical shell nearly rhombic, smooth, one and a half times as long as broad, with four strong, conical, radial spines, two on the poles of the principal axis (as prolongations of the sagittal septa), two others on the poles of the transverse axis. Transverse girdle narrow, with two pores on the isthmus. Sagittal septa longer than the medullary shell. Eight gates triangular.

_Dimensions._--Length of the medullary shell 0.04, breadth 0.03; length of the cortical shell 0.18, breadth 0.12.

_Habitat._--Tropical Atlantic, Station 347, depth 2250 fathoms.

{652}7. _Octopyle tetrastyle_, n. sp.

Cortical shell lentelliptical, with rough surface and four angular radial spines in the lateral plane, opposite in pairs in two crossed diagonals. Lateral girdle nearly rectangular (each angle with one spine), one and a third times as long as broad. Sagittal septum longer than the hexagonal medullary shell. Eight gates egg-shaped.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.03; length of the cortical shell 0.16, breadth 0.12.

_Habitat._--North Atlantic, Station 354, surface.

8. _Octopyle tetraptera_, n. sp.

Cortical shell quadrangular, with thorny surface and four latticed wing-like prolongations on the corners of the quadrangle, opposite in pairs in two crossed diagonals, each wing supported by a strong radial spine. Lateral girdle quadrangular, one and a half times as long as broad. On the isthmus of the transverse girdle four pores. Sagittal septum longer than the hexagonal medullary shell. Eight gates triangular.

_Dimensions._--Length of the medullary shell 0.07, breadth 0.04; length of the cortical shell 0.24, breadth 0.16.

_Habitat._--Pacific, central area, Station 271, surface.

9. _Octopyle stenozona_, n. sp. (Pl. 9, fig. 11).

Cortical shell quadrangular, with thorny surface, and four conical spines in the lateral plane, on the points, where both cortical girdles are crossed. Lateral girdle very narrow, nearly square, with rounded corners, of equal length and breadth. On the isthmus of the narrow transverse girdle only one pore. Sagittal septum of the same length as the subspherical medullary shell. Eight gates irregular roundish or nearly circular.

_Dimensions._--Diameter of the medullary shell 0.04, of the cortical shell 0.14.

_Habitat._--Tropical Atlantic, Station 338, depth 1990 fathoms.

10. _Octopyle euryzona_, n. sp.

Cortical shell nearly quadrangular, very thorny, with four bunches of stronger radial spines on the four corners. Lateral girdle broad, with rounded edges of the quadrangle, little longer than broad. On the isthmus of the broad transverse girdle four pores. Sagittal septum of the same length as the elliptical medullary shell, with many radial beams. Eight gates nearly square.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.035; length of the cortical shell 0.16, breadth 0.14.

_Habitat._--Southern Atlantic, Station 332, depth 2200 fathoms.

{653}11. _Octopyle hexastyle_, n. sp.

Cortical shell nearly rectangular, thorny, with six strong radial spines in the lateral plane, two opposite on the poles of the principal axis (prolongations of the principal beam at the sagittal septum), four others on the truncate corners of the quadrangular lateral girdle, which is one and a half times as long as broad. On the isthmus of the broad transverse girdle three pores. Sagittal septum longer than the hexagonal medullary shell. Eight egg-shaped gates.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.04; length of the cortical shell 0.18, breadth 0.12.

_Habitat._--Northern Atlantic, surface; Canary Islands, Station 9, depth 3150 fathoms.

12. _Octopyle sexangulata_, n. sp. (Pl. 9, fig. 12).

Cortical shell thorny, hexagonal, with six strong conical radial spines on the six corners of the lateral girdle, lying in the lateral plane, two opposite on the poles of the transverse axis, four others in pairs on both sides of the annular sagittal constriction, which is only twice as long as the elliptical medullary shell. Transverse girdle broad, with five pores on the isthmus. Eight gates roundish, smaller than the medullary shell.

_Dimensions._--Length of the medullary shell 0.045, breadth 0.035; length of the cortical shell 0.11, breadth 0.13.

_Habitat._--Pacific, central area, Station 270, surface.

13. _Octopyle octostyle_, n. sp.

Cortical shell elliptical, thorny, with eight long cylindrical radial spines, lying in two crossed diagonal planes, opposite in pairs, quite as in the nearly related _Tetrapyle octacantha_. It differs from this species by the perfect, latticed, vertical sagittal septum, which divides every gate into two triangular halves. Lateral girdle elliptical, one and a third times as long as broad. Transverse girdle on the isthmus with three pores. Septum longer than the elliptical medullary shell.

_Dimensions._--Length of the medullary shell 0.06, breadth 0.045; length of the cortical shell 0.16, breadth 0.12.

_Habitat._--South Pacific, Station 288, surface.

14. _Octopyle obtecta_, n. sp.

Cortical shell quadrangular, thorny, with eight strong diagonal spines, lying in two crossed planes, and arising from the proximal edges of the gates in the same way as in the foregoing species. It differs from this by four prominent latticed roofs, which arise from the distal edge of the gates and connect the diagonal spines of one gate. _Octopyle obtecta_ bears therefore the same relation to _Octopyle octostyle_ that _Tetrapyle pluteus_ does to _Tetrapyle octacantha_.

_Dimensions._--Length of the medullary shell 0.055, breadth 0.04; length of the cortical shell 0.16, breadth 0.12.

_Habitat._--Indian Ocean, surface, Madagascar, Rabbe.

{654}15. _Octopyle decastyle_, n. sp. (Pl. 9, fig. 13).

Cortical shell thorny, nearly quadrangular, with ten stronger radial spines, two opposite on the poles of the principal axis (as prolongations of the axial beams), eight others opposite in pairs in two diagonal planes (as in the foregoing species). Lateral girdle nearly square, with slightly convex lateral faces, slightly concave principal faces. Transverse girdle broad, with four pores on the isthmus. Sagittal septum of the same length as the hexagonal medullary shell. Eight gates elliptical or irregular roundish.

_Dimensions._--Length of the medullary shell 0.05, breadth 0.032; diameter of the cortical shell 0.15.

_Habitat._--Western Tropical Pacific, Station 224, depth 1850 fathoms.

Genus 285. _Pylonium_,[324] Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Pylonida# with trizonal lentelliptical medullary shell surrounded by three crossed latticed cortical girdles, one smaller (primary) transverse, one larger (secondary) lateral, and one (tertiary) sagittal girdle.

The genus _Pylonium_ is the most highly developed among the Diplozonaria, with six complete latticed girdles, three on the medullary shell, three on the cortical shell, the latter corresponding to the former. In each of the two systems the transverse girdle is smaller than the lateral; the sagittal girdle may be smaller or larger than the lateral girdle. _Pylonium_ repeats the typical form of _Trizonium_; but whilst in this latter the medullary centre of the shell is a simple spherical chamber, here in _Pylonium_ it is a true trizonal or _Larnacilla_-shaped medullary shell.

Subgenus 1. _Pylonissa_, Haeckel.

_Definition._--Cortical shell smooth or thorny, but without large, symmetrically disposed spines.

1. _Pylonium circozonium_, n. sp.

Cortical shell thorny, without radial spines; all its three girdles nearly of the same size, subcircular, very narrow, only with one to two rows of pores. Four gates subcircular.

_Dimensions._--Principal axis 0.12, transverse axis 0.11; diameter of the subspherical medullary shell 0.05.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

{655}2. _Pylonium hexazonium_, n. sp.

Cortical shell smooth, without symmetrical radial spines. Transverse girdle elliptical, one and a half times as long as broad. Lateral and sagittal girdles nearly circular. Four gates egg-shaped.

_Dimensions._--Principal axis 0.16, transverse axis 0.15; diameter of the subspherical medullary shell 0.04.

_Habitat._--South Pacific, Station 295, depth 1500 fathoms.

3. _Pylonium nephropylium_, n. sp.

Cortical shell thorny, without radial spines. All its three girdles elliptical, one and a third times as long as broad. Four gates kidney-shaped, nearly twice as broad as high.

_Dimensions._--Principal axis 0.17, transverse axis 0.13; length of the lentelliptical medullary shell 0.04, breadth 0.03.

_Habitat._--Pacific, central area, Station 273, depth 2350 fathoms.

Subgenus 2. _Pylonura_, Haeckel.

_Definition._--Cortical shell armed with large, symmetrically disposed spines.

4. _Pylonium quadricorne_, n. sp. (Pl. 9, fig. 14).

Cortical shell thorny, with four strong horn-like curved spines in the lateral plane. All its three girdles elliptical, one and a third times as long as broad. Four gates kidney-shaped, twice as broad as high. (This species is a further developmental stage of _Tetrapyle quadricornis_, with perfect sagittal girdle.)

_Dimensions._--Principal axis 0.16, transverse axis 0.12; length of the medullary shell 0.05, breadth 0.04.

_Habitat._--Pacific, central area, Stations 270 to 274, surface; Indian Ocean, Madagascar, Rabbe, surface.

5. _Pylonium octacanthum_, n. sp.

Cortical shell thorny, with eight long and thin cylindrical radial spines, lying in two crossed diagonal planes, opposite in pairs, and arising from the eight points where the transverse girdle is crossed by the lateral girdle. All three girdles elliptical, one and a half times as long as broad; four gates triangular. (This species is a further developmental stage of _Tetrapyle octacantha_, with perfect sagittal girdle.)

_Dimensions._--Principal axis 0.18, transverse axis 0.13; length of the medullary shell 0.03 to 0.04.

_Habitat._--Cosmopolitan; Mediterranean, Atlantic, Pacific, surface.

{656}6. _Pylonium stenozonium_, n. sp.

Cortical shell thorny, with eight short conical radial spines, lying in two crossed diagonal planes (as in the foregoing species). All three girdles nearly of the same size, subcircular, very narrow, only with one to two rows of pores. Four gates kidney-shaped.

_Dimensions._--Principal axis 0.15, transverse axis 0.14; diameter of the quadrangular medullary shell 0.04.

_Habitat._--Equatorial Atlantic, Station 347, surface.

Subfamily 3. TRIPLOZONARIA, Haeckel.

_Definition._--#Pylonida# with three concentric systems of fenestrated girdles, lying in three concentric lentelliptical faces (every system with one to three girdles lying in one lentelliptical face).

Genus 286. _Amphipylonium_,[325] Haeckel, 1881, Prodromus, p. 463.

_Definition._--#Pylonida# with trizonal lentelliptical medullary shell, surrounded by a double latticed cortical shell; inner cortical shell _Pylonium_-shaped, with three perfect crossed girdles; outer cortical shell only represented by a single (transverse) girdle.

The genus _Amphipylonium_ opens the series of Triplozonaria, or of those Pylonida in which the shell is composed of three concentric systems of latticed girdles. The first (and innermost) system represents the complete trizonal medullary shell, which is probably a lentelliptical _Larnacilla_-shell. The first and intermediate system is formed of a complete trizonal cortical shell of the same form, but much larger (like _Pylonium_). The third (and outermost) system is represented by one to three latticed girdles, corresponding to the former and forming an outer or second cortical shell. In _Amphipylonium_ (as the most simple form of the Triplozonaria) there is only developed the first (transverse) girdle of the third system. It repeats therefore the form of _Amphipyle_, the cortical shell of which is here double (Prodromus, 1881, p. 463).

1. _Amphipylonium semilunare_, n. sp.

Inner cortical shell lentelliptical, one and a half times as long as broad, with smooth surface and four semilunar gates. Transverse girdle of the outer cortical shell with two smooth semilunar wings, with convex, smooth lateral crests; each wing as broad as the transverse girdle of the inner cortical shell and twice as long as its lateral girdle.

_Dimensions._--Length of the lentelliptical medullary shell 0.04, breadth 0.03; length of the inner cortical shell 0.15, breadth 0.1; length of each lateral wing of the outer cortical shell 0.3, breadth 0.1.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

{657}2. _Amphipylonium spinosissimum_, n. sp.

Inner cortical shell lentelliptical, one and a third times as long as broad, with thorny surface and four kidney-shaped gates. Transverse girdle of the outer cortical shell very spiny, with two semilunar wings, with convex thorny lateral crests; each wing one and a half times as broad as the transverse girdle of the inner cortical shell, and one and a half times as long as its lateral girdle.

_Dimensions._--Length of the lentelliptical medullary shell 0.05, breadth 0.03; length of the inner cortical shell 0.16, breadth 0.12; length of each lateral wing of the outer cortical shell 0.24, breadth 0.18.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

3. _Amphipylonium tetraceros_, n. sp.

Inner cortical shell hexagonal, one and a half times as long as broad, with smooth surface and four transverse elliptical gates. Transverse girdle of the outer cortical shell smooth, with rectilinear smooth lateral crests, which are prolonged at both ends into strong conical spines (therefore four spines in the lateral plane); each wing of the same breadth as the transverse girdle of the inner cortical shell, and twice as long as its lateral girdle.

_Dimensions._--Length of the hexagonal medullary shell 0.03, breadth 0.02; length of the inner cortical shell 0.12, breadth 0.08; length of each lateral wing of the outer cortical shell 0.24, breadth 0.08.

_Habitat._--Northern Pacific, Station 253, depth 3125 fathoms.

4. _Amphipylonium octoceros_, n. sp.

Inner cortical shell quadrangular, one and a third times as long as broad, with thorny surface and four kidney-shaped gates. Transverse girdle of the outer cortical shell thorny, constricted at the equator, with concave thorny lateral crests, and with eight strong radial spines, opposite in pairs and lying in two crossed diagonal planes; each wing narrower than the transverse girdle of the inner cortical shell and twice as long as its lateral girdle.

_Dimensions._--Length of the subspherical medullary shell 0.04, breadth 0.035; length of the inner cortical shell 0.14, breadth 0.11; length of each lateral wing of the outer cortical shell 0.26, breadth 0.09.

_Habitat._--Southern Pacific, Station 295, depth 1500 fathoms.

Genus 267. _Tetrapylonium_,[326] Haeckel, 1881, Prodromus, p. 464.

_Definition._--#Pylonida# with trizonal lentelliptical medullary shell, surrounded by a double latticed cortical shell; inner cortical shell _Pylonium_-shaped, with three perfect crossed girdles; outer cortical shell represented by two crossed girdles, one (smaller) transverse and one (larger) lateral girdle.

{658}The genus _Tetrapylonium_ represents a further developmental stage of the foregoing _Amphipylonium_; whilst in this latter the outer cortical shell (or the third system of girdles) is formed only by a transverse girdle, here this is crossed by a lateral girdle. _Tetrapylonium_ repeats therefore the typical form of _Tetrapyle_, but with doubled cortical shell (Prodromus, 1881, p. 464).

1. _Tetrapylonium pantellipticum_, n. sp.

Outer cortical shell elliptical, one and a third times as long as broad, with smooth surface and four elliptical gates. Inner cortical shell elliptical, with smooth surface and four elliptical gates which repeat the form of the four outer gates, but are of half the size. Transverse girdle broad, with five pores on the isthmus.

_Dimensions._--Length (or principal axis) of the first shell (medullary shell) 0.04, breadth (or transverse axis) 0.03; length of the second (or inner cortical) shell 0.12, breadth 0.09; length of the third (or outer cortical) shell 0.18, breadth 0.14.

_Habitat._--Pacific, central area, Station 274, depth 2750 fathoms.

2. _Tetrapylonium reniforme_, n. sp.

Outer cortical shell elliptical, one and a half times as long as broad, with thorny surface and four large kidney-shaped gates. Inner cortical shell of the same form, but one-third smaller, also with four kidney-shaped gates. Transverse girdle small, but with two pores on the isthmus.

_Dimensions._--Length of the first (innermost) shell 0.03, breadth 0.02; length of the second (middle) shell 0.14, breadth 0.08; length of the third (outermost) shell 0.2, breadth 0.11.

_Habitat._--Indian Ocean, Zanzibar, Pullen, depth 2200 fathoms.

3. _Tetrapylonium quadrangulare_, n. sp. (Pl. 9, fig. 15).

Outer cortical shell nearly quadrangular, one and a third times as long as broad, with four rounded corners, from which arise in the lateral plane four three-sided pyramidal radial spines, opposite in pairs in two crossed diagonal axes. Surface thorny; four gates elliptical or nearly quadrangular. Inner cortical shell more elliptical, by one half smaller, with broader transverse girdle and four kidney-shaped gates. Medullary shell nearly spherical, of half the size.

_Dimensions._--Length of the first shell 0.045, breadth 0.035; length of the second shell 0.1, breadth 0.07; length of the third shell 0.16, breadth 0.11.

_Habitat._--South Atlantic, off Tristan da Cunha, Station 332, depth 2200 fathoms.

4. _Tetrapylonium octacanthum_, n. sp.

Outer cortical shell elliptical, one and a third times as long as broad, with thorny surface, and eight long and thin radial spines arising from the corners of the four triangular gates, and lying {659}opposite in pairs in two crossed diagonal planes. Inner cortical shell of the same shape, but two-thirds smaller, also thorny. Transverse girdle broad, with four pores on the isthmus. (This species appears to be the common _Tetrapyle octacantha_, with doubled cortical shell.)

_Dimensions._--Length of the first shell 0.04, breadth 0.03; length of the second shell 0.16, breadth 0.12; length of the third shell 0.24, breadth 0.18.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

5. _Tetrapylonium armatum_, n. sp.

Outer cortical shell nearly quadrangular, one and a fourth times as long as broad, with very spiny surface. Between numerous smaller (simple or branched) thorns arise twelve larger radial spines, symmetrically distributed, eight wing-spines crossed in two diagonal planes (as in _Tetrapylonium octacanthum_) and four corner spines on the four rounded corners of the lateral plane (as in _Tetrapylonium quadrangulare_). Therefore this species combines the armature of both foregoing species. Four gates kidney-shaped. Inner cortical shell elliptical, smooth, three-fourths smaller.

_Dimensions._--Length of the first shell 0.045, breadth 0.035; length of the second shell 0.18, breadth 0.15; length of the third shell 0.25, breadth 0.2.

_Habitat._--Pacific, central area, Station 266, depth 2750 fathoms.

Genus 288. _Pylozonium_,[327] n. gen.

_Definition._--#Pylonida# with trizonal lentelliptical medullary shell, surrounded by double latticed cortical shell; inner cortical shell as well as the outer _Pylonium_-shaped, each with three perfect crossed girdles (transverse, lateral, and sagittal).

The genus _Pylozonium_ is the most highly developed form of the Pylonida, as all three systems of concentric girdles here become fully developed, each system with three perfect girdles. In this genus only among all the Pylonida we find nine complete latticed girdles, and in each of the three dimensive planes three concentric elliptical girdles. The first system forms the medullary shell, the second system the inner cortical shell, and the third system the outer cortical shell.

1. _Pylozonium novemcinctum_, n. sp.

Outer cortical shell lentelliptical, thorny, one and a third times as long as broad, without radial spines, with four kidney-shaped gates. Inner cortical shell of the same shape, but one-third smaller, four times as large as the lentelliptical medullary shell.

_Dimensions._--Length of the medullary shell 0.04, breadth 0.03; length of the inner cortical shell 0.16, breadth 0.12; length of the outer cortical shell 0.24, breadth 0.18.

_Habitat._--North Pacific, Station 253, depth 3125 fathoms.

{660}2. _Pylozonium octacanthum_, n. sp. (Pl. 9, fig. 16).

Outer cortical shell lentelliptical, one and a third times as long as broad, with thorny surface and eight long and thin radial spines arising from the corners of the four elliptical gates, and lying opposite in pairs in two crossed diagonal planes. Inner cortical shell of the same shape but one-fourth smaller, about twice as large as the lentelliptical medullary shell.

_Dimensions._--Length of the medullary shell 0.065, breadth 0.045; length of the inner cortical shell 0.18, breadth 0.12; length of the outer cortical shell 0.24, breadth 0.18.

_Habitat._--North Pacific, Station 244, depth 2900 fathoms.

Family XXVII. #THOLONIDA#, n. fam. (Pl. 10).

_Definition._--#Larcoidea# with regular, completely latticed cortical shell, which is composed of two to six or more hemispherical or cap-shaped domes (vaulted chambers or cupolas). The domes lie opposite in pairs on the poles of the three dimensive axes, are separated by annular constrictions, and surround a simple or _Larnacilla_-shaped central chamber.

The family #Tholonida# represents a peculiar and very remarkable group of the #Larcoidea#, distinguished from the other groups of this suborder by the characteristic form of the shell, composed of a variable number of hemispherical domes or cupolas. The middle and original part of the shell is constantly formed of an elliptical or subspherical central chamber, which often, but not constantly, contains a small medullary shell. An even number (two, four, six, or more) of domes is attached to the poles of the three dimensive axes of the central chamber; according as only one, or two, or all three axes develop cupolas, we distinguish in this family three different subfamilies (the Amphitholida, Staurotholida, and Cubotholida). The Amphitholida (or Tholonida monaxonia) form cupolas only on the two poles of one single axis, and this axis corresponds to the minor (or transverse) axis of the central chamber, we find here therefore constantly at least two lateral cupolas (Pl. 10, figs. 1-7). The Staurotholida (or Tholonida diaxonia) form cupolas on the poles of two axes perpendicular one to another; these two axes are the major (principal) and the minor (transverse) axis of the central chamber, we find here therefore constantly at least four cupolas crossed in pairs (Pl. 10, figs. 8-11). The Cubotholida (or Tholonida triaxonia) form cupolas on the poles of all three dimensive axes (perpendicular one to another); corresponding to the principal, transverse, and sagittal axes of the lentelliptical central chamber; therefore we find here constantly at least six cupolas, attached in pairs on the six sides of the central chamber (Pl. 10, figs. 12-17).

The number of genera and species in the family #Tholonida# is rather large, but the number of individuals is much smaller than in most of the other SPUMELLARIA, and {661}particularly than in the nearly allied Pylonida; the greater number of the species are very rare. Besides this they seem to be very variable and inclined to produce numerous abnormalities. Very often intermediate forms of transition are to be found between the Tholonida and other #Larcoidea#, particularly the Pylonida and Lithelida. In many species the thick-walled shell is very opaque, and offers great difficulties to the clear study; in the greater number the structure of the shell cannot be understood completely without rolling the shell to the different sides; and the distinction between the Staurotholida and Cubotholida is often very difficult.

The primordial chamber of the Tholonida, or the central chamber in which its growth begins, is either a simple lentelliptical shell (without enclosed medullary shell), like _Cenolarcus_, or it is a trizonal shell, like _Larnacilla_, and contains a small concentric medullary shell. But this important inner shell of the central chamber offers peculiar difficulties for study. In many cases (probably in the greater number) there is no doubt the same characteristic trizonal medullary shell, which we found in the Larnacida and Pylonida, and this is our principal argument, if we regard the Tholonida as #Larcoidea#, which are most nearly allied to both these families, and in which the fenestrated open cortical girdles of the Pylonida are replaced by fenestrated closed cupolas or domes; the characteristic "gates" or large fissures in the cortical shell of the former are therefore here perfectly closed by network. The Tholonida agree in this point with the Larnacida, but are distinguished from them by the prominent vaultings of the hemispherical cupolas or domes, which give them a peculiar appearance. Each pair of domes (opposite on both poles of one dimensive axis) corresponds to one single girdle of the Pylonida.

Regarding the absence or presence of a medullary shell in the central chamber, we may divide the Tholonida into two groups, Cenotholida (without medullary shell) and Coccotholida (with medullary shell). The absence of the medullary shell in many Tholonida may be either primary (original) or secondary (by reduction and loss of it). Probably in the larger proportion (if not always?) its absence is the consequence of reduction and loss, and in this case the Cenotholida must be derived phylogenetically from the Coccotholida, but possibly often (or always?) the contrary may also be the case. At present we cannot find certain arguments for one opinion or the other.

The connection between the medullary (internal) and the cortical (external) shell of the central chamber in the Coccotholida is effected by two lateral wings of a latticed transverse girdle; on both sides of the latter remain the four internal open "gates" of the Pylonida and Larnacida. Very often the limiting edges of these four gates are prolonged into eight external radial spines which lie opposite in pairs in two crossed diagonal planes, and correspond to the eight portal-spines of _Tetrapyle octacantha_. Besides these, we often find four other beams opposite in pairs in the two crossed axes (principal and lateral). Very often also these radial beams (between medullary and {662}cortical shells) are prolonged on the outside into prominent free radial spines. But other spines may also arise from the surface. Very remarkable is the presence of twenty symmetrically disposed radial spines in some forms of Amphitholida (_Amphitholus acanthometra_ and _Amphitholonium acanthonium_, &c.). It recalls the twenty radial spines of the #Acanthonida#, though the laws of distribution in the two similar cases are quite different.

The cortical shell of the Tholonida remains either simple, or it becomes doubled by formation of an outer veil or envelope. This outer cortical shell or "veil" may either repeat completely the typical form of the inner with its cupolas, or it may form an ellipsoidal envelope without dome-shaped partitions. Its network is either like that of the inner, or it is very delicate and irregular, spider-web like. The connection between the two cortical shells is effected by a variable number of radial beams, often prolonged on the outside into radial spines.

The growth of the Tholonida by apposition of new chambers is very characteristic, constantly pairs of chambers originating at both opposite poles of the three dimensive axes, firstly on the transverse axis, secondly on the principal axis, thirdly on the sagittal axis. It is possible that by repeated apposition of new chambers the Tholonida are transformed into #Discoidea#, but commonly the number of domes is restricted, two in the Amphitholida, four in the Staurotholida, six in the Coccotholida. If new chambers in an irregular manner or in a spiral order be opposed to those first formed pairs of chambers, the Tholonida may be transformed into Soreumida, Streblemida, or Lithelida. The phylogenetic connection between these families is probably very complex and very worthy of further research.

The network of the shell in the Tholonida is sometimes regular, with circular (often hexagonally framed) pores of equal size, sometimes irregular, with roundish pores of unequal size. Commonly the shell is thick and compact; in those genera in which the cortical shell is doubled, the inner is commonly compact, the outer a very delicate veil.

The central capsule of the Tholonida is originally always enclosed by the central chamber, and like this of lentelliptical form, a triaxial ellipsoid. With the apposition of cupolas the growing central capsule may form dome-shaped protuberances which enter into the former. So we may find in the Amphitholida a central capsule with three joints (separated by two annular parallel constrictions), in the Staurotholida a cross-shaped capsule (with four caps around the central lentellipsis), in the Cubotholida a clustered capsule with six caps, surrounding six sides of the cubical central mass. In the Coccotholida (with medullary shell in the central chamber) the latter encloses in the centre the nucleus of the cell. The calymma, or the jelly-veil between the central capsule and the enveloping shell, probably always exhibits annular constrictions, corresponding to those which separate the cupolas of the shell. All these anatomical peculiarities in the Tholonida require a further accurate study.

{663}_Synopsis of the Genera of Tholonida._

-------------------------------------------------------------------------
I. Subfamily Amphitholida.
Cupolas developed only in the direction of one single axis (commonly
two cupolas).
(Tholonida unidimensiva.)
-------------------------------------------------------------------------
{Two simple cupolas
{ (without veil), 289. _Tholartus_.
Central chamber simple, {
without medullary shell. {Two double cupolas
{ (with veil), 290. _Tholodes_.

{Two simple cupolas
Central chamber { (without veil), 291. _Amphitholus_.
_Larnacilla_-shaped, with {
enclosed medullary shell. {Two double cupolas
{ (with veil), 292. _Amphitholonium_.
-------------------------------------------------------------------------
II. Subfamily Staurotholida.
Cupolas developed in the direction of two axes perpendicular one to
another (commonly four cupolas).
(Tholonida bidimensiva.)
-------------------------------------------------------------------------
{Four simple cupolas
{ (without veil), 293. _Tholostaurus_.
Central chamber simple, {
without medullary shell. {Four double cupolas
{ (with veil), 294. _Tholoma_.

{Four simple cupolas
Central chamber { (without veil), 295. _Staurotholus_.
_Larnacilla_-shaped, with {
enclosed medullary shell. {Four double cupolas
{ (with veil), 296. _Staurotholonium_.
-------------------------------------------------------------------------
III. Subfamily Cubotholida.
Cupolas developed in the direction of three axes perpendicular one
to another (commonly six cupolas).
(Tholonida tridimensiva.)
-------------------------------------------------------------------------
{Six simple cupolas
{ (without veil), 297. _Tholocubus_.
Central chamber simple, {
without medullary shell. {Six double cupolas
{ (with veil), 298. _Tholonium_.

{Six simple cupolas
Central chamber { (without veil), 299. _Cubotholus_.
_Larnacilla_-shaped, with {
enclosed medullary shell. {Six double cupolas
{ (with veil), 300. _Cubotholonium_.

Subfamily 1. AMPHITHOLIDA, Haeckel.

_Definition._--#Tholonida# with monaxial growth; cupolas opposite on the poles of one axis. (Shell commonly three-chambered, with two domes on both sides of the central chamber.)

Genus 289. _Tholartus_,[328] n. gen.

_Definition._--#Tholonida# with simple cortical shell (without external veil); with two hemispherical cupolas, opposite on the poles of one axis; central chamber between them simple (without medullary shell).

{664}The genus _Tholartus_ (Pl. 10, fig. 1) represents the most simple form of all Tholonida, a three-jointed simple cortical shell without medullary shell; two hemispherical cupolas or domes are attached on both lateral sides of a simple triaxial ellipsoidal central chamber, corresponding to the poles of its lateral or transverse axis. _Tholartus_ may be regarded as the common ancestral form of all Tholonida, and may be derived either from the Pylonida _Monozonium_ (by complete lattice-locking of the lateral wings) or from the Ellipsida _Cenellipsis_ (by development of two lateral protuberances on the poles of the transverse axis). But it is also possible that the absence of the medullary shell is produced by reduction, and in this case _Tholartus_ may be descended from _Amphitholus_.

Subgenus 1. _Tholartella_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial spines.

1. _Tholartus tricolus_, n. sp. (Pl. 10, fig. 1).

Central chamber one and a half times as high and as broad as both cupolas. Surface of the shell rough. Pores regular, circular, without hexagonal frames, three times as broad as the bars; about sixteen on the half meridian of the central chamber (or its vertical diameter).

_Dimensions._--Length of the shell (major axis of the central chamber, vertical) 0.1, breadth of the shell (major axis of the whole three-chambered shell, horizontal) 0.13; pores 0.01, bars 0.003.

_Habitat._--Pacific, central area, Station 270, depth 2925 fathoms.

2. _Tholartus paniscus_, n. sp.

Central chamber of the same breadth, but of the double height of both cupolas. Surface of the shell rough. Pores regular, circular, with hexagonal frames, of the same breadth as the bars; eight to ten on the half meridian of the central chamber.

_Dimensions._--Length of the shell (major axis of the central chamber, vertical) 0.12, breadth of the shell (major axis of the whole three-chambered shell, horizontal) 0.15; pores and bars 0.008.

_Habitat._--Pacific, central area, Station 271, depth 2425 fathoms.

3. _Tholartus isocolus_, n. sp.

Central chamber of the same breadth and height as both cupolas. Surface of the shell smooth. Pores irregular, roundish, once to three times as broad as the bars; ten to fifteen on the half meridian of the central chamber.

_Dimensions._--Length of the shell 0.11, breadth 0.14; pores 0.004 to 0.01, bars 0.003.

_Habitat._--Pacific, central area, Station 270, depth 2925 fathoms.

{665}Subgenus 2. _Tholartissa_, Haeckel.

_Definition._--Surface of the shell with radial spines.

4. _Tholartus tripanis_, n. sp.

Central chamber of the same size as both cupolas. Surface of the shell thorny, everywhere covered with short conical radial spines, about as long as the cupolas. Pores regular or subregular, circular, twice as broad as the bars; eight to ten on the half meridian.

_Dimensions._--Length of the shell 0.1, breadth 0.12; pores 0.01, bars 0.005.

_Habitat._--South Pacific, Station 295, surface.

5. _Tholartus sagitta_, n. sp.

Central chamber twice as large as both cupolas. Surface of the shell spiny, with eight to twelve regularly (?) distributed radial spines, radiating from the two constrictions between the three chambers. Spines needle-shaped, very thin and long. Pores subregular or irregular, roundish or circular, about three times as broad as the bars; twelve to sixteen on the half meridian.

_Dimensions._--Length of the shell 0.11, breadth 0.15; pores 0.01, bars 0.003.

_Habitat._--North Pacific, Station 244, surface.

Genus 290. _Tholodes_,[329] n. gen.

_Definition._--#Tholonida# with double cortical shell (with external veil), with two hemispherical cupolas, opposite on the poles of one axis; central chamber between them simple (without medullary shell).

The genus _Tholodes_ (Pl. 10, fig. 2) differs from _Tholartus_, its probable ancestral form, only in the duplication of the shell; the outer shell has exactly the same three-jointed form as the inner; both are connected by eight radial beams, lying in two diagonal planes and corresponding to the eight frontal spines of _Tetrapyle octacantha_. Possibly _Tholodes_ may also be descended from _Amphitholonium_ by loss of the medullary shell. I have observed only one single specimen of this genus.

1. _Tholodes cupula_, n. sp. (Pl. 10, fig. 2).

Outer shell of the same form and structure as the inner shell. Central chamber larger than both cupolas. Surface a little rough, without radial spines. Distance of both shells equals the height of the outer cupolas. Pores of both shells regular, circular, three times as broad as the {666}bars; six to seven in the semicircle of one inner cupola, ten to twelve in the semicircle of one outer cupola. Both shells are connected by eight radial beams, crossed in two diagonal planes.

_Dimensions._--Major axis of the outer shell 0.1, minor 0.065; major axis of the inner shell 0.05, minor 0.04; pores 0.006, bars 0.002.

_Habitat._--Western Tropical Pacific, Station 225, depth 4475 fathoms.

Genus 291. _Amphitholus_,[330] n. gen.

_Definition._--#Tholonida# with simple cortical shell (without external veil), with two hemispherical cupolas, opposite on the poles of one axis; central chamber (between them) _Larnacilla_-shaped, with medullary shell.

The genus _Amphitholus_ is the most simple form of all Coccotholida (or all Tholonida provided with medullary shell). Its simple cortical shell exhibits, like _Tholartus_, two lateral hemispheric cupolas, attached on both sides of a lentelliptical central chamber (on the poles of its transverse axis). The central chamber has the form of a triaxial ellipsoid, and contains, like _Larnacilla_, a small medullary shell, connected with it by the wings of a latticed transverse girdle. _Amphitholus_ may be derived either from _Tholartus_ by secondary production of a cortical shell, or (more probably) from _Larnacilla_ by apposition of two lateral dome-shaped protuberances. In the latter case it may be regarded as an _Amphipyle_, the lateral open wings of which are closed by lattice-work.

Subgenus 1. _Amphitholissa_, Haeckel.

_Definition._--Surface of the shell smooth or rough, without radial spines.

1. _Amphitholus artiscus_, n. sp. (Pl. 10, fig. 3).

Central chamber of the same height as both cupolas. Surface of the cortical shell rough. Pores regular, circular, with prominent hexagonal frames, twice as broad as the bars; eight to ten in the basal semicircle of one cupola.

_Dimensions._--Major axis of the shell 0.16, minor axis 0.1; pores 0.008, bars 0.004.

_Habitat._--Pacific, central area, Station 272, depth 2600 fathoms.

2. _Amphitholus artidium_, n. sp.

Central chamber one-fourth higher than both cupolas. Surface of the cortical shell smooth. Pores irregular, roundish, once to three times as broad as the bars. (Very similar to _Tholartus_ tricolus, Pl. 10, fig. 1, but different by the medullary shell.)

_Dimensions._--Major axis of the shell 0.15, minor axis 0.12; pores 0.004 to 0.01, bars 0.003.

_Habitat._--North Pacific, Station 241, depth 2300 fathoms.

{667}Subgenus 2. _Amphitholura_, Haeckel.

_Definition._--Surface of the shell with radial spines or thorns.

3. _Amphitholus octacanthus_, n. sp.

Central chamber about twice as high and broad as both cupolas. From the surface arise, symmetrically distributed, eight stout conical radial spines, lying in two diagonal planes, as prolongations of the eight wing-beams, which connect the _Tetrapyle_-shaped medullary shell with the two ring-like constrictions of the cortical shell. Pores of the latter regular, circular, twice as broad as the bars; ten to twelve in the basal semicircle of one cupola.

_Dimensions._--Major axis of the shell 0.15, minor 0.12; pores 0.01, bars 0.005.

_Habitat._--South Atlantic, Station 332, depth 2200 fathoms.

4. _Amphitholus dodecanthus_, n. sp.