Chapter L: M N is a guide for the eye, in order to direct it in the axis of the (13)
The hydra pallens has been fully described only by M. Rösel;[113] it is very seldom to be met with, is of a pale yellow colour, and grows smaller gradually from the bottom, the tail is somewhat round or knobbed, the arms are about the length of the body, of a white colour, and generally seven in number, apparently composed of a chain of globules; it brings forth the young from all parts of its body. Linnæus defines it as, hydra pallens tentaculis subsenis mediocribus;[114] Pallas as, hydra attenuata corpore flavescente, sursum attenuato.[115]
[113] Insecten Belustigung, 3. Theil. pag. 465. Tab. LXXVI. LXXVII.
[114] System. Nat. p. 1320, No. 4.
[115] Zoophyt. 4.
OF THE HYDRA HYDATULA.
Plate XXI. Fig. 1, 2, 3, and 4.
The next in order is the hydra hydatula, which we have already defined from Linnæus as a hydra with four obsolete arms, and a vesicular body: it is spoken of by several medical writers, who are enumerated in the Systema Naturæ, p. 1321. It is described also by Hartman, Misc. Nat. Cur. Dec. I. An. 7, Obs. 206, Dec. II. An. 4, Obs. 73, as hydatis animata; also in the Dissert. de Inf. Viv. p. 50; n. 6, tænia hydatoidea. Pallas defines it as tænia hydatigena rugis imbricata corpore postice bulla lymphaticæ terminato. The following description is extracted from that in the Philosophical Transactions, No. 193, by Dr. Tyson, who names it lumbricus hydropicus.
In the dissection of a gazella or antelope, Dr. Tyson observed several hydatides or films filled with water, about the size of a pigeon’s egg, and of an oval form, fastened to the omentum, and some in the pelvis, between the bladder of urine and the rectum; and he then suspected them to be a particular sort of insect, bred in animal bodies, or at least the embryos or eggs of them: 1. Because he observed them included in a membrane, like a matrix, so loosely, that by opening it with a finger or knife, the internal bladder, containing the serum or lympha, seemed no where to have any connection with it, but would very readily drop out, still retaining its liquor, without spilling any of it. 2. He observed that this internal bladder had a neck or white body, more opake than the rest of the bladder, and protuberant from it, with an orifice at its extremity, by which, as with a mouth, it exhausted the serum from the external membrane, and so supplied its bladder or stomach. 3. Upon bringing this neck near the candle, it moved and shortened itself. Fig. 1, represents one of these watery bladders inclosed in its external membrane, its shape was nearly round, being only a little depressed or flatted, as a drop of quicksilver will be by lying on a plane. In Fig. 2, the neck is better seen; the external membrane being taken off, an open orifice is found at its extremity; it consists of circular rings or incisures, which are more visible when magnified, as in Fig. 3; it then appears granulated with a number of little eminences all over the surface; the orifice at the extremity seems to be formed by retracting itself inwards, and upon trial it was found to be so; for in Fig. 4, the neck of this polype is represented magnified and drawn out its whole length; on opening it there were found within the two strings a, a, which probably convey into the stomach the moisture and nourishment, which the animal, by protruding its neck, extracts from the external membrane.[116]
[116] Hydra hydatula habitat in abdomine mammalium, ovium, suum,
murium, &c. inter peritoneum et intestina. Vesica lymphatica,
pellucida, magnitudine pruni, petiolata corpore cylindrico, in cujus
apice os, quod, corpore compresso, movet tentacula vix manifesta.
Linn. Syst. Nat. p. 1321, No.5.
OF THE HYDRA STENTOREA.
Plate XXII. Fig. 27 and 28.
Hydra tentaculis ciliaribus corpore infundibuliformi.
The arms of this hydra are rows of short hairs, the body trumpet-shaped.
This species of hydra is very common, and has been described by almost every writer on these subjects; it is placed by Müller among the vorticellæ.
Vorticella stentorea caudata, elongata, tubæformis limbo ciliato. Müller animalcula infusoria.
Mr. Baker originally named it the funnel-like polype, which Messrs. Trembley and Reaumur changed to the tunnel-like polype, under which name it appears in the Philosophical Transactions, No. 474.
There are three kinds of them, which are of different colours, green, blue, and white. The white ones are the most common. It is necessary to observe them often, and in various attitudes, in order to obtain a tolerable idea of their structure. They do not form clusters, but adhere singly by their tail to whatever comes in their way; their anterior end is wider than the posterior, and being round, gives the animal somewhat of a funnel form, though it is not completely circular, having a sort of slit or gap that interrupts the circle. The edge of this opening is furnished with a great number of fibrillæ, which by their brisk and continual motions excite a current of water; the small bodies that float or swim near this current, are forced by it into the mouth of the little animal. Trembley says, that he has often seen a number of very small animalcula fall one after another into the mouth, some of which were afterwards let out again at another opening, which he was not able to describe.
They can fashion their mouths into several different forms. If any thing touch them, they shrink back and contract themselves. They live independent of each other, swimming freely through the water in search of their prey, and fix to any thing they meet with.
These animals multiply by dividing themselves, not longitudinally, nor transversely, but sloping and diagonal wise; the proceedings in nature continually varying in every new form of life. Of the two polypes produced by the division of one, the first has the old head and a new tail; the other, the old tail and a new head.
To make the description more clear, Trembley called that with the old head the superior polype, that with the new head the inferior one. The first particular that is observable in these polypes, when they are going to divide, is the lips of the inferior one; a transverse and oblique stripe indicates the part where it is going to divide; the new lips are formed at about two-thirds of the length of the polype, reckoning from the head; the division is made in a sloping line, that goes about half way round the parent animal; these lips are at first discerned by a slow motion, which engages the attention of the observer. They then insensibly approach each other and close, whereby a swelling is formed on the side of the polype, which is soon found to be a new head. When the swelling is considerably increased, the two polypes may be plainly distinguished. The superior one being now connected with the inferior one only by its lower extremity, is soon detached from it, and swims away to fix itself on some convenient substance; the inferior one remains fastened to the place where the original polype was fixed before the division.
From the various modes by which different species of polypes are multiplied, we are led to form more exalted ideas of nature, and to see that the little we discover is but an exceeding small part of her contents; we learn also to be more cautious in reasoning from analogy, and laying down the known for a model to the unknown, because we find that the operations in nature are varied ad infinitum.
The growth of the hydra fusca is very quick, but that of the hydra stentorea is much more so. The progress of the fœtus is always more rapid than that of the infant and adult animal; but in these organized atoms the evolution is so rapid, as to appear almost like an immediate creation.
Fig. 28 represents the hydræ stentoreæ, or funnel-polypes, fixed to the under side of a piece of some vegetable substance; they are in this figure of their natural size.
Fig. 27, the same polypes magnified; the different forms they assume are also seen here, sometimes short and thick, as at m m; long, as at n; nearly globular, as at o; extended to the full size, as at k; seen as contracted at i. The fibrillæ or little hairs may be seen in most of the attitudes except those of l.
OF THE HYDRA SOCIALIS.
Plate XXI. Fig. 11.
Hydra socialis mutica torosa rugosa.[117]
[117] Linn. Syst. Nat. p. 1321. No. 7.
Social hydra, bearded thick and wrinkled.
This species of hydra has been described by many writers. It is the vorticella socialis of Müller, who defines it as vort cella caudata, aggregata, clavata; disco obliquo. Müller Animalcula Infusoria, p. 304. Pallas makes it a brachionus, Pall. Zooph. 53.
In Fig. 11, these animals are represented as considerably magnified; they appear like a circle, surrounded with crowns, or ciliated heads, tied by small thin tails to a common center, from whence they advance towards the circumference, where they turn like a wheel, with a great deal of vivacity and swiftness, till they occasion a kind of whirlpool, which brings into its sphere the proper food for the polype. When one of them has been in motion for a time, it stops, and another begins; sometimes two or three may be perceived in motion together. They are often to be found separate, with the tail sticking in the mud. The body contracts and dilates very much, so as sometimes to have the appearance of a cudgel; at others, to assume almost a globular form. The young polypes of this species have been sometimes taken for the hydra stentorea.
OF THE VORTICELLÆ.
We now come to another division of these animals, to which later writers have given the name of vorticellæ; this term I shall therefore adopt, being of opinion that it behoves every man to maintain that order in scientific arrangement which is not inconsistent with truth, except he can produce another arrangement more expressive of the nature of the objects it is designed to discriminate; a process requiring no small degree of attention.
The variety that may be observed in these minute animals confirms a principle, which, the more it is inquired into, the more it will be found to accord with the general operations in nature, namely, that there is always a pre-existent principle of life necessary to the organization both of animals and vegetables; that the alimentary and other particles which are added to, or apparently belong to them, produce nothing of themselves; they are incapable of forming the least fibre, but they are able to become constituent parts of one organical whole, together with the instruments whereby the forming principle is manifested, and rendered capable of acting upon certain orders of creatures.
VORTICELLA.
Animal calyce vasculoso; ore contractili ciliato, terminali. Stirps fixa.
A small animal, with a vascular cup; the mouth is at one end ciliated, and capable of being contracted, the stem fixed.
VORTICELLA ANASTATICA.
Plate XXI. Fig. 13, 14, 15, and 16*.
Vorticella anastatica, composita, floribus campanulatis, stirpe multiflora rigescente.
Vorticella anastatica, compound, with bell-shaped flowers, and a rigid stem.
Cluster polype, second species. Trembley, Philos. Trans. vol. xliv. part. 2. p. 643.
These polypes form a group resembling a cluster, or more properly an open flower; this flower or cluster is supported by a stem, which is fixed by its lower extremity to some of the aquatic plants or extraneous bodies that are found in the water; the upper extremity forms itself into eight or nine lateral branches, perfectly similar to each other; these have also subordinate branches, whose collective form much resembles that of a leaf. Every one of these assemblages is composed of one principal branch or nerve, which makes with the main stem of the cluster an angle somewhat greater than a right one; from both sides of this nerve the smaller lateral branches proceed; these are shorter the nearer their origin is to the principal branch.
At the extremity of the principal branch, and also of all the lateral ones, there is a polype or vorticella. There are others on both sides of the lateral twigs, but at different distances from their extremity. These polypes are all exceeding small, and of a bell-like figure; near their mouth a quick motion may be discerned, though not with sufficient distinctness to convey an adequate idea of its cause; upon the branches of these clusters are round bodies, which will be more particularly described presently.
Every cluster has eight or nine of these branches or leaves; they do not all proceed from the same point, but the points from whence they set out are not far asunder; each of these branches is bent a little inwards, so that all of them taken together form a kind of shallow cup. If the eye be placed right over the base of this cup, the appearance of the whole eight or nine branches is like unto that of a star, with so many rays proceeding from the center. If the cluster be slightly touched, all the branches instantly fold up, and form a small round mass. The stem which supports the cluster contracts also at the same time, folding up like a workman’s measuring rule, that consists of three or four joints. This extraordinary assemblage constitutes one organized whole, formed of a multitude of similar and particular ones. A new species of society, in which all the individuals are members of each other in the strictest sense, and all participate of the same life.
A few days after one of these clusters is formed, small round bodies or bulbs may be perceived to protrude in several places from the body of the branch; these grow very fast, and arrive at their greatest growth in two or three days. The bulbs detach themselves from the branches out of which they spring, and go away, swimming till they can settle upon some substance which they meet with in the water, and to which they fix themselves by a short pedicle; the bulbs are then round, only a little flatted on the under side, the pedicle continues to lengthen gradually for about twenty-four hours, during the same time the bulbs also change their figure, and become nearly oval. There are in a cluster but few of these bulbs, compared with the number of the vorticellæ, neither do all the bulbs come out at the same time. The bulb then divides lengthways into two smaller ones, but which are still much larger than the vorticellæ themselves. It is not long before these are separated like the first, and thus form four bulbs on the same stalk; these again divide themselves, and form eight; which again subdivide, and consequently make sixteen. They are all connected with the stalk by a proper pedicle, but they are not all of an equal size; the largest continue to divide, and the smallest begin to open, and take the bell-formed shape. Trembley observed from one round bulb, in about twenty-four hours, by repeated divisions, one-hundred and ten vorticellæ to be formed.
It has been asked with propriety, what plant or what animal could have led us to expect an existence and mode of propagation similar to that of the vorticella anastatica?
Fig. 13 represents one branch of the vorticella anastatica; on this branch, besides the vorticellæ which are of a bell-like form, some of those round bodies from which they first spring, and by which they are so remarkably distinguished from any other species, may be seen.
Fig. 14 represents one of the globular bodies after it has parted from the cluster, and has fixed itself to some other body, and after the globule itself and its pedicle have begun to lengthen.
Fig. 15 represents the two bodies that were formed by the parting of that which is shewn at Fig. 14.
Fig. 16* represents four that were formed by the separation of the two bulbs, exhibited in the foregoing figure.
VORTICELLA PYRARIA.
Plate XXII. Fig. 25, 26.
Vorticella composita, floribus muticis obovatis; tentaculis bigeminis, stirpe ramosa. Compound, with beardless oval florets, two double arms, the stem branched.
It is somewhat of a pear shape, the base is pellucid, the top truncated, the lateral arms, which are a pair on each side, cannot be distinguished without some attention; they are sometimes to be seen disengaged from the pedicle, and rolling swiftly in a kind of circle.
VORTICELLA CRATÆGARIA.
Plate XXII. Fig. 40.
Vorticella composita, floribus muticis globosis; tentaculis binis, stirpe ramosa. Compound, with globous naked florets, two tentacules, and a branched stem.
These vorticellæ are to be found in the month of April, both in the mud, and upon the tail of the monoculus quadricornis; they are generally heaped together in the manner in which they are represented in the figure; they are of a spherical form, and united to one common stalk. They are also often to be found without any pedicle. The body is rather contracted; the aperture is circular, and surrounded with a marked margin; it has two small arms. With a deep magnifier, a vehement rotatory motion may be seen. They sometimes separate from the community, and go forwards in a kind of spiral line, and then in a little time come back again to the rest.
The figure represents a parcel of these vorticellæ united together.
Among the other authorities for this animal, Linnæus refers to Baker’s description of the mulberry insect, “Employment for the Microscope,” p. 348, which, as it differs a little from the preceding account, we shall insert here. That from which his drawing was made, and which he has described, was found in a ditch near Norwich; he called it the mulberry insect, from the resemblance it bore to that fruit; though the protuberances that stand out round it are more globular than those of a mulberry. It is to be seen rolling about from one place to another, and is probably a congeries of animalcula; they are to be met with in different numbers of knobs or protuberances, some having fifty or sixty, others more or less, down to four or five. The manner of moving is the same in all. They are generally of a pale yellow.
VORTICELLA OPERCULARIA.
Plate XXII. Fig. 29.
Vorticella composita floribus muticis ovalibus, stirpe ramosa. Compound, with naked oval florets, and a branched stem.
These vorticellæ are of a lemon shape, and are generally found in clusters, branching out from a stem, which mostly adheres to some convenient substance.
That species of them which is described by Baker had a very short pedicle, and the animals were much longer than those which are represented at Fig. 29. There was no main stem, but all the pedicles were joined in one center, round which the animals extended themselves as so many radii, forming a very pleasing figure.
The mouths of these animalcula are not ciliated, but they are furnished with a round operculum or cover, connected by a long ligament or muscle, which extends downwards through the body, and is affixed withinside of it, near the tail. This ligament may be contracted or dilated, so that the cover can be removed to some distance from the mouth; in this situation several short hairs maybe found to radiate from it; these have a vibratory motion, by which they excite a current of water, most probably to draw in the proper nourishment, after which they shut or pull down the cover, which they again extend at pleasure: when the cover is pulled close down, the mouth contracts, and no hairs are to be seen.
Fig. 29 represents the vorticella opercularia; ſ, the operculum removed at some distance from the mouth, at t; it is nearly close at r, the mouth contracted, the cover drawn in, and no hairs to be seen; u, a part of the stalk, from which some of the animalcula are separated.
VORTICELLA UMBELLARIA.
Plate XXII. Fig. 30.
Vorticella composita, floribus ciliatis globosis muticis, stirpe umbellata. Compound, with ciliated globous naked florets and an umbellated stem.
Vorticella acinosa, simplex, globosa, granis nigricantibus, pedunculo rigido. Müller Animal. Infus. p. 319.
We frequently find in divers places, upon water-plants, and other bodies in the water, a whitish substance that looks like mould; plants, pieces of wood, snail shells, &c. are often entirely covered over with this substance. If we examine any of these minute bodies by the microscope, we shall find such motions as will induce us to think them an assemblage of living animals, severally fixed to the extremities of small stems or pedicles, many of which are often so united as to form together a sort of branches or clusters, from whence they have been termed clustering polypes, or des polypes en bouquet.
These clusters are larger or smaller, according to the species of the vorticellæ which form them, as well as owing to the concurrence of many other circumstances. To obtain a clear idea of the figure of these animals, it is best to observe the smaller clusters, as in the larger they are often rendered less distinct on account of the number.
The length of those which are represented at Fig. 30, is about the 240th of an inch; they are of a bell-shape. The anterior part a c generally appears open, the posterior part is fixed to a stem or pedicle, b e; it is by the extremity of this pedicle that the vorticella fastens itself to any substance. It appears in the microscope of a brownish colour, excepting at the smaller end b, where it is transparent, as well as the whole pedicle b e. When the anterior part a c is open, a very lively motion may be perceived about its edges; and when it presents itself in a particular manner, something very much resembling the little wheels of a mill, moving with great velocity, may be discovered on both sides of the edges of this anterior part.
These vorticellæ are able to contract themselves suddenly. They may be made to do this, either by touching them, or moving the substance to which they are fixed. When they contract, the edges of the anterior parts are drawn quite into the body; on resuming their former posture, the edges may be seen to come forth, and put themselves in motion as before. Minute substances that float in the water are often forced down into these openings, and sometimes are thrown out again.
They are capable of swimming about singly, but their form is in that case considerably different from that which they have when they are fixed. To see regularly in what manner the clusters are formed, and in what way these little creatures multiply, it is best to observe one that is fixed by itself.
The pedicle of a single vorticella is at first short, but it soon grows longer, and then begins to multiply, that is, to divide or split itself into two lengthways. To effect this, the lips are first drawn into the body, the anterior part closes and becomes round, and loses its bell shape, the motion about the lips ceases, though a small degree of motion may be perceived within the body. The anterior end flattens gradually, and spreads wider in proportion as it grows smaller. It then gradually splits down the middle, that is, from the middle of the head to the pedicle, so that in a little time two separate round bodies appear to be joined to the end of the pedicle that before supported but one.
The mouth or anterior part of each of these bodies now opens by degrees; and in proportion as they open, the lips of the new vorticella begin to display themselves. The motion before spoken of may then also be perceived. Indeed it is the best time of observing it; it is at first slow, but more rapid in proportion as the mouth opens, when it is as swift as that of the vorticella before it began to divide, and we may now look upon it as completely formed. A vorticella is generally about one hour in dividing itself.
The lower of the three drawings, Fig. 30, represents two vorticellæ joined by their posterior extremity to one pedicle; soon after the division, each vorticella begins to shew a pedicle of its own.
Fig. 30 represents a cluster of eight vorticellæ; by this figure we may form some idea in what manner the pedicles are disposed as their number increases. There were at first only two at b, whose branches lengthened to d, and then each of them was divided into two, now forming four; these again lengthened and reached i; when they were again subdivided, as in the figure.
The reader will join with Bonnet in admiring the group of wonders afforded by a single spot of mouldiness. What unforeseen, varied, and interesting scenes are presented within so small a compass! what a theatre is exhibited to a thinking mind! But our abode is so recluse, that we have but a glimmering view of it: how great would our astonishment be, if the whole spectacle was disclosed to us at once, and we were enabled to penetrate into the interior structure of this wonderful assemblage of living atoms! Our eyes see only the gross parts of the decorations, whilst the machines that execute them remain in impenetrable darkness! Who shall enlighten this profound obscurity, or dive into an abyss where reason is lost; or draw from thence the treasures of wisdom concealed within it? Let us learn to be content with the small portion that is communicated to us, and contemplate with gratitude the first traces of human understanding that are imparted to us in these discoveries.
VORTICELLA BERBERINA.
Vorticella composita, floribus ovalibus muticis, stirpe ramosa. Compound, with oval beardless florets.
This is a species of the vorticellæ, which resembles the preceding one in many respects, particularly in being multiplied in the same manner, that is, by dividing or splitting, according to its length.
They are more slender than the vorticella umbellaria; the branches of the clusters are transparent. When many of them are together, they appear of a changeable violet colour; the clusters are not unlike a sprig of spun glass. The motion of the lips is not so easily distinguished as in the foregoing species, though it may be observed in these whilst they are opening and completing their formation. For at these times the motion is but slow, whereas it becomes afterwards very quick in those that are arrived at a state of perfection.
All the cluster vorticellæ detach themselves from time to time from the stem, and from these they swim about till they fix again upon some convenient substance; the branches, when deserted, bear no more vorticellæ.
VORTICELLA DIGITALIS.
Plate XXII. Fig. 31.
Vorticella composita, floribus cylindricis, unisulcatis semiclausis, stirpe ramosa. Compound, with cylindrical florets.
Vorticella composita, cylindrica, crystallina, apice truncata et fissa, pedunculo fistuloso ramosa. Müller Animal. Infus. p. 327.
This species of the vorticella is very scarce, it seems only to have been seen by Rösel, who found it on the monoculus quadricornis, till it was discovered in the year 1784 by Müller, who had sought for it several years before, but in vain.
The body is cylindrical, crystalline, and appears almost empty; it has three pellucid points disposed lengthways, the apex is truncated in an oblique direction, the margin bent back. The upper part contracts itself, and the margin then assumes a conical shape, with a convex surface; there are in general but few branches from the principal stem, and these are short and thick. It excites an undulatory motion, but no hairs, nor any rotatory motion, have been discovered. Fig. 31, o and n, represents the vorticella adhering to the monoculus quadricornis.
VORTICELLA CONVALLARIA.
Plate XXII. Fig. 39.
Vorticella simplex, gregaria, flore campanulata mutico; tentaculis bigeminis, stirpe fixa. Simple, but gregarious, the florets bell-shaped, with two pair of little arms, and a fixed stem.
Vorticella simplex, campanulata, pedunculo rotortili. Müller Animal. Infus.
These vorticellæ, or bell-animals, as they are termed by Baker, are generally found adhering to some substance in the water; they are represented here as found by Rösel, fixed to a curious cornu ammonis, with points projecting from the back. To the naked eye they appear only as so many little white points, but under the microscope, as little bells, agitating the water to a considerable distance. The stems of these have a particular motion, they draw themselves up and shorten all at once, taking the form of a spiral wire or screw; in a moment after they again resume their former shape, stretching themselves out straight as before. Many of them may be seen at times adhering to each other by their tails; the cilia, which are two on each side of the mouths, are very seldom to be perceived.
VORTICELLA URCEOLARIS.
Plate XXII. Fig. 33, 34, 35, 36, 37, 38.
Vorticella simplex, pedunculata, ore dentato. Single, with a short tail, and toothed mouth.
Brachionus capsularis testa ovata apice sexdentata basi incisa, cauda longa bicuspi. Müller Animal. Infus. p. 356.
To the naked eye it appears as a white moveable point; but when examined by the microscope, a tail projecting from the lower part is discovered, and a double rotatory instrument is seen, which it can conceal or expose at pleasure. It has been seen and described by most microscopical writers; but as Baker’s seems to be the most perfect description, I shall principally follow his account of it.
He discovered three species of them, two of which are included under the vorticella urceolaris. Fig. 33, 34, 35, are of the first species; Fig. 36, 37, 38, are of the second kind. The first sort, when extended, is about twice as long as it is broad. It is contained in a shell; the fore part of this is armed with four sharp teeth or points; the opposite side has no teeth, but is waved or bent in two places, like the form of a Turkish bow. At the bottom there is a hole, through which it pushes the tail. It fastens itself by this tail to any convenient substance when it intends to use its rotatory organs; but when it is floating in the water, and at all other times when not adhering to any body, it wags the tail backwards and forwards something like a dog.
We may consider it as divided into a head, thorax, and abdomen; each of which may be extended and contracted considerably: it can, by dilating all three, protrude the head beyond the shell, or by contracting them, draw the whole body within the same.
The head, when extended, divides itself into two branches, between which, another part, a kind of proboscis, is pushed out; at the end of this are two fibrils, that appear when they are at rest like a broad point, but which can be moved to and from each other very briskly with a vibratory motion, see Fig. 33.
The form and situation of the two branches are sometimes changed, the ends thereof becoming more round, and the vibratory motion is altered to a rotatory one: this alteration is represented at Fig. 34: the head also appears in this figure. The thorax is annexed to the lower part of the head; it is muscular: within it there is a moving intestine, which has been supposed to be either the lungs or the heart of the little creature, see b, Fig. 33 and 34.
A communication is formed between the thorax and the abdomen by means of a short vessel c, whose alternate contractions and dilatations occasion the abdomen to rise and fall alternately, having at the same time a sort of peristaltic motion. The food is conveyed through this vessel into the abdomen, where it is digested; it is then discharged by the anus, which is placed near the tail.
The tail has three joints, and is cleft or divided at the extremity, by which means it can better fasten itself to suitable objects. It is in general projected from the lower end of the shell, moving nimbly to and fro, serving the animal as a rudder when it is swimming, to direct its course.
When the water in which the little animal is placed is nearly dried away, or when it has a mind to compose itself to rest, it contracts the head and fore-part of the body, brings them down into the shell, and pulls the tail upwards, so that the whole of this minute creature is contained within the shell, see Fig. 35. The shell is so transparent that the terminations cannot be easily distinguished when the animal is extended; but whatever is transacted within the shell, is as plain as if there was no substance between the eye and the interior parts.
Fig. 36, 37, 38, exhibit the appearance of another species of these animals, which differs from the foregoing kind. This has also a head, a thorax, and abdomen, but then they are not separated by a gut or intermediate vessel, as in the former, but are joined immediately together, and at the place where in the first kind a moveable intestine was seen; in this a muscle, most probably the heart, may be discovered; it has a regular systole and diastole: this part is intended to be shewn at a, Fig. 36, 37, 38. Like the other, it draws the head and tail within the shell, which then appears to have six teeth or spikes on one side, and two on the other. It very seldom protrudes its head so far out as the other; sometimes the fibrillæ may be seen within the margin of the shell.
Both species carry their young in an oval integument or bag, fastened externally to the lower part of the shell, somewhere about the tail; these bags are sometimes opake at one end, and seemingly empty at the other, see d, Fig. 34: sometimes the middle is opake, with a transparent margin, see b, Fig. 36.
It is highly entertaining to see a young one burst its integument, and gradually force its way out; in performing this operation, it is much assisted by the motion of the tail of the parent. The head part comes out first, it then sets its rotatory organ in motion, by which it is completely disengaged, leaving the integument behind, which the vorticella freed itself from by repeated strokes with its tail. A young one almost disengaged is seen at b, Fig. 38; another embryo, c, was left adhering to the shell.
There are four more species of the vorticellæ mentioned by Linnæus, which are, the vorticella encrinus, the vorticella polypina, the vorticella stellata, and the vorticella ovifera; which, being marine animals, do not come properly within our plan. The vorticella polypina will be described hereafter.
TUBULARIA CAMPANULATA.
Plate XXII. Fig. 32.
Tubularia reptans, tubis campanulatis. Creeping, with campanulated tubes.
It is called by Baker the bell-flowered, or plumed animal.
These little creatures dwell in colonies together, from ten to fifteen in number, living in a kind of slimy mucilaginous case, which, when expanded in the water, has some resemblance to a bell with its mouth upwards. These bells or colonies are to be found adhering to the large leaves of duck-weed and other aquatic plants.
The bell or case which these animals inhabit, being very transparent, all the motions of its inhabitants may be discerned distinctly through it. There are several ramifications or smaller bells proceeding from the larger one; in each of these there is an inhabitant. The opening at the top of these bells is just large enough for the creature’s head, and a small part of its body to be thrust out from it, the rest remaining in the case, into which it also draws the head on the least alarm.
Besides the particular and separate motions which each of these creatures is able to exert within its case, and independent of the rest, the whole colony has a power of altering the position of the bell, and removing it from one place to another. These animalcula seem not to like to dwell in societies, whose number exceeds fifteen; when the colony happens to increase in number, the bell may be observed to split gradually, beginning from about the middle of the upper extremity, and proceeding downwards towards the bottom, till they at last separate and become two colonies, independent of each other.
The arms are very near each other; sixty may often be counted in one plume, having each the figure of an Italic _ʃ_, one of whose hooked ends is fastened to the head; and altogether, when expanded, compose a figure somewhat like a horseshoe, convex on the side next the body, but gradually opening and turning outwards, so as to leave a considerable distance within the outer extremities of the arms.
The plumed polype is of a very voracious disposition, devouring a great number of small animals. If the arms, when extended, be observed attentively with the microscope, they will be found to have a constant vibratory motion; alternately bending withinside of the plume, and then rising up again. When one arm ceases its motion, the same is performed by another; thus by the perpetual agitation of the several arms, such a strong current is produced in the water, as brings the animalcula, and other minute bodies, that are floating near the polype, into its mouth, which is situated between the arms. The food, if agreeable to the creature, is swallowed; if otherwise, it is rejected by a contrary motion.
The animal may be seen very plain when it has retired within the tube. The body is about one-eighth of an inch long, without reckoning the plume, which is about the same length. It is cylindrical, and the skin is very transparent. The plume is only a continuation of this transparent skin, it is very broad in proportion to the body, and of a remarkable figure; the base is of the shape of a horseshoe; from this base the arms project, they bend rather outwards. The plume which they form, gives them a resemblance to some flowers. The arms may be compared, from their fineness and transparency, to very fine threads of glass. The base of the plume is grooved, and is fixed to the animal by the middle of the horseshoe which it forms, and it is here that there is an opening which serves as a mouth to the animal. The intestines are easily distinguished through its transparent skin; when it has just been eating, they are of a deep brown colour. Three principal parts are very visible, the oesophagus, the stomach, and the rectum.
In the inside of these animals a small oblong whitish body is formed, which is carried to the outside, and remains fixed in a perpendicular direction to the body; many of these are formed daily, and of these oval bodies new animals are produced, exactly similar to the parent.
If these minute bodies be eggs, they are of a singular kind, being destitute of any covering, and are neither membranaceous nor crustaceous; we cannot with propriety say the young ones are hatched from them; we can, however, perceive these oviform bodies to unfold themselves gradually. The developement is accomplished in a few minutes, and an animalculum appears like the parent.
Trembley amassed a great number of these eggs, and carried them from England with him, keeping them quite dry; on putting them into water, they gradually developed, and became as perfect as the tubularia from which they proceeded.
There is a very great similarity in the construction of this little creature and many of the marine polypes, who, like it, exist in tubes of the same growth with themselves.
Fig. 32 represents three tubulariæ campanulatæ or plumed polypes very much magnified, namely, one, b f a c d d e h g i, which is out of its cell; e h, the oesophagus; f g, the stomach; a f, the rectum; a c d d e, the plume, consisting of the base a e, which is but little seen, and the arms c d d, which proceed from the edges of this base; a second polype, _A B I_, which is within its cell, and in which the skin containing the plume is reversed. The third polype, s t u u, is a young one exhibited out of its cell; g o o, threads which are fixed at one end to the intestines of the animal, by the other to the bottom of the cell, l k.
CHAP. VIII.
OF THE ANIMALCULA INFUSORIA.
Our knowledge of the microscopic world is at present very contracted, but we know enough to give us high conceptions of its concealed wonders, and to fill us with profound astonishment at the infinite variety of forms that are made recipient of life. A few of the inhabitants of this minute world have been discovered. The figure and apparent habits of life of these, resemble so little those with which we are more acquainted, that it is often difficult to find terms to express what is represented to the eye.
Animalculum signifies a little animal, and therefore the term might be applied to every animal which is considerably inferior in size to ourselves. It has been customary, however, to distinguish by the name of animalcula only such animals as are of a size so diminutive, that their true figure cannot be discerned without the assistance of glasses; and more especially it is applied to such as are altogether invisible to the naked eye, and cannot even be perceived to exist, but by the aid of microscopes.
By the help of magnifying glasses we are brought into a kind of new world; and numberless animals are discovered, which, from their minuteness, must otherwise for ever have escaped our observation: and how many kinds of these invisibles there may be, is yet unknown; as they are observed of all sizes, from those which are barely invisible to the naked eye, to such as resist the action of the microscope, as the fixed stars do that of the telescope, and with the best magnifiers hitherto invented, appear only as so many moving points.
The smallest living creatures our instruments can shew, are those that inhabit the waters; for, though possibly animalcula equally minute, or perhaps more so, may fly in the air, or creep upon the earth, it is scarce possible to obtain a view of them; whereas, water being transparent, and confining the creatures within it, we are enabled, by applying a drop of it to our glasses, to discover with ease a great part of its contents, and in a space barely visible to the naked eye, often perceive a thousand little creatures, all full of life and vigour.
By the animalcula infusoria are meant, not the larvæ of those insects which in their first state are inhabitants of water, and afterwards become winged insects, as the gnat, &c. Baker, and many other writers on the subject, have often confounded these, and hence entered into a train of reasoning contrary to fact and experience. The animalcula infusoria take their name from their being found in all kinds either of vegetable or animal infusions; if seeds, herbs, or other vegetable substances, be infused in water, it will soon be filled with an indefinite number of these minute beings. There is a prodigious variety in their forms; some perfectly resemble the bell-polype; others are round or oblong, without any, at least apparent, members; some resemble a bulb with a long taper tail; some are nearly spherical; the greater part are vesicular and transparent. Those most generally found in every drop of ditch water are mere inflated bladders, with a small trace of intestines in the center; the next are a flat kind, with a number of legs under the belly.
Motion seems to be their great delight; they pervade with equal ease and rapidity, and in all forms and directions, the whole dimensions of the drop, in which they find ample space for their various progressions, sometimes darting straight forward, at other times moving obliquely, then again circularly: they know how to avoid with dexterity any obstacles that might obstruct their progress. Hundreds may be seen in a drop of water in constant action, yet never striking against each other. If at any time the clusters prove so thick as to impede any of their motions, they roll and tumble themselves over head, creep under the whole range, force their way through the midst, or wheel round the cluster, with surprizing swiftness; sometimes they will suddenly change the direction in which they are moving, and take one diametrically opposite thereto. By inclining the glass on which the drop of water is laid, it may be made to move in any direction; the animalcula in the drop will swim as easily against the stream as with it.
If the water begin to evaporate, and the drop to grow smaller, they flock impetuously towards the remaining part of the fluid; an anxious desire of attaining this momentary respite of life is very visible, as well as an uncommon agitation of the organs by which they imbibe the water. These motions grow more languid as the water fails, till at last they entirely cease.
Animalcula and insects will support a great degree of cold, but both one and the other perish when it is carried beyond a certain point. The same degree of heat that destroys the existence of insects, is fatal to animalcula; as there are animalcula produced in water at the freezing point, so there are insects which live in snow.
If the smallest drop of urine be put into a drop of water where these animalcula are roving about, apparently happy and easy, they instantly fly to the other side, but the acid soon communicating itself to this part, their struggles to escape are increased, but the evil also increasing, they are thrown into convulsions, and soon expire.
Among animalcula, as in every other part of nature, there is constantly a certain proportion preserved between the size of the individuals and their number. There are always fewest amongst the larger kinds, but they increase in number as they diminish in size, till of the last, or lowest to which our powers of magnifying will reach, there are myriads to one of the larger. Like other animals, they increase in size from their birth till they have attained their full growth. When deprived of food, they grow thin and perish; and different degrees of organization are to be discovered in their structure.
The birth and propagation of these microscopic beings is as regular as that of the largest animals of our globe; for though their extreme minuteness prevents us, in most cases, from seeing the germ from which they spring, yet we are well assured, from numerous observations, that the manner in which they multiply is regulated by constant and invariable laws.
It has been shewn that different species of the hydræ and vorticellæ multiply and increase by natural divisions and subdivisions of the parent’s body; this manner of propagation is very common among the animalcula in infusions, though with many remarkable varieties. Some multiply by a transverse division, a contraction takes place in the middle, forming a kind of neck that becomes smaller every instant, till they are enabled by a slight degree of motion to separate from each other. These animalcula in general studiously avoid each other; but when they are in the labour of multiplication, and the division is in great forwardness, it is not uncommon to see one of them precipitate itself on the neck of the dividing animalculum, and thus accelerate the separation.
Another species, when it is on the point of multiplying, fixes itself to the bottom of the infusion; it then forms an oblong figure, afterwards becomes round, and begins to turn rapidly, as if upon an internal center, continually changing the direction of its rotatory motion; after some time, we may perceive two lines on the spherule, forming a kind of cross; soon after which the animalculum divides into four distinct beings, which grow, and are again subdivided.
Some multiply by a longitudinal division, which in one kind begins in the fore-part, and others in the hind-part; from another kind a small fragment is seen to detach itself, which very soon acquires the form of the parent animalculum. Lastly, some propagate in the same manner as those we deem more perfect animals.
From what has been said, it appears clearly that their motions are not purely mechanical, but are produced by an internal spontaneous principle, and that they must therefore be placed among the class of living animals, for they possess the strongest marks, and the most decided characters of animation; and consequently, that there is no foundation for the supposition of a chaotic and neutral kingdom, which can only have derived its origin from a very transient and superficial view of these animalcula.
It may also be further observed, that as we see the motions of the limbs, &c. of the more noble animals, viz. the human species, are produced by the mechanical construction of the body and the action of the soul thereon, and are forced by the ocular demonstration arising from anatomical dissection, to acknowledge this mechanism which is adapted to produce the various motions necessary to the animal; and as when we have recourse to the microscope, we find those pieces which had appeared to the naked eye as the primary mechanical causes of the particular motions, to consist themselves of lesser parts, which are the causes of motion, extension, &c. in the larger; when the structure can therefore be traced no further by the eye or glasses, we have no right to conclude, that the parts which are invisible, are not equally the subject of mechanism: for this would be only to assert in other words, that a thing may exist because we see and feel it, and has no existence when it is not the object of our senses.
The same train of reasoning may be applied to microscopic insects and animalcula; we see them move, but because the muscles and members which occasion these motions are invisible, shall we infer that they have not muscles, with organs appropriated to the motion of the whole and its parts? To say that they exist not, because we cannot perceive them, would surely not be a rational conclusion. Our senses are indeed given us, that we may comprehend some effects; but then we have also a mind with reason bestowed upon us, that from the things which we do perceive with our senses, we may deduce the nature of those causes and effects which are imperceptible to the corporeal eye.
Messrs. Buffon, Needham, and Baron Münchhausen, have considered this part of animated nature in so different a light from other writers, that we cannot with propriety entirely pass them over. Needham imagined that there was a vegetative force in every microscopical point of water, and every visible filament of which the whole vegetable contexture consists; that the several species of microscopic animals may subside, resolve again into gelatinous filaments, and again give lesser animals, and so on, till they can be no further pursued by glasses. That agreeable to this idea, every animal or vegetable substance advances as fast as it can in its revolution, to return by a slow descent to one common principle, whence its atoms may return again, and ascend to a new life. That notwithstanding this, the specific seed of one animal can never give another of a different species, on account of the preparation it must receive to constitute it this specific seed.
Buffon asserts, that what have been called spermatic animals, are not creatures really possessing life, but something proper to compose a living creature, distinguishing them by the name of organic particles, and that the moving bodies which are to be found in the infusions either of animal or vegetable substances, are of the same nature.
Baron Münchhausen supposed that the seeds of mushrooms were first animals, and then vegetables; and this, because he had observed some of the globules in the infusions of mushrooms, after moving some time, to begin to vegetate.
It might be sufficient in the first instance to observe, that Messrs. Needham, and Buffon, by having recourse to a vegetative force and organic particles, to account for the existence and explain the nature of animalcula, and the difficulties of generation, have substituted words in the place of things; and that we are no gainers by the substitution, unless they explain the nature of these powers. But to this we may add, that all those who have examined the subject with accuracy and attention, as Bonnet, De Saussure, Baker, Wrisberg, Spalanzani, Haller, Ellis, Müller, Ledermüller, Corti, Rofredi, &c. disagree with the foregoing gentlemen, proving that they had deceived themselves by inaccurate experiments, and that one of them, Buffon, had not seen the spermatic animals he supposed himself to be describing, insomuch that Needham was at last induced to give up his favourite hypothesis.
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