Chapter III: Part 3

EGERIA, an ancient Italian goddess of springs. Two distinct localities were regarded as sacred to her,--the grove of Diana Nemorensis at Aricia, and a spring in the immediate neighbourhood of Rome at the Porta Capena. She derives her chief importance from her legendary connexion with King Numa, who had frequent interviews with her and consulted her in regard to his religious legislation (Livy i. 19; Juvenal iii. 12). These meetings took place on the spot where the sacred shield had fallen from heaven, and here Numa dedicated a grove to the Camenae, like Egeria deities of springs. After the death of Numa, Egeria was said to have fled into the grove of Aricia, where she was changed into a spring for having interrupted the rites of Diana by her lamentations (Ovid, _Metam._ xv. 479). At Aricia there was also a Manius Egerius, a male counterpart of Egeria. Her connexion with Diana Nemorensis, herself a birth goddess, is confirmed by the fact that her aid was invoked by pregnant women. She also possessed the gift of prophecy; and the statement (Dion. Halic. ii. 60) that she was one of the Muses is due to her connexion with the Camenae, whose worship was displaced by them.

EGERTON, SIR PHILIP DE MALPAS GREY, Bart. (1806-1881), English palaeontologist, was born on the 13th of November 1806, the son of the 9th baronet. He was educated at Eton and Christ Church, Oxford, where he graduated B.A. in 1828. While at college his interest in geology was aroused by the lectures of W. Buckland, and by his acquaintance with W.D. Conybeare. Subsequently when travelling in Switzerland with Lord Cole (afterwards 3rd earl of Enniskillen) they were introduced to Prof. L. Agassiz at Neufchatel, and determined to make a special study of fossil fishes. During the course of fifty years they gradually gathered together two of the largest and finest of private collections--that of Sir Philip Grey Egerton being at Oulton Park, Tarporley, Cheshire. He described the structure and affinities of numerous species in the publications of the Geological Society of London, the _Geological Magazine_ and the Decades of the Geological Survey; and in recognition of his services the Wollaston medal was awarded to him in 1873 by the Geological Society. He was elected F.R.S. in 1831, and was a trustee of the British Museum. As a member of Parliament he represented the city of Chester in 1830, the southern division of Cheshire from 1835 until 1868, and the western division from 1868 to 1881. He died in London on the 6th of April 1881. His collection of fossil fishes is now in the British Museum.

EGG, AUGUSTUS LEOPOLD (1816-1863), English painter, was born on the 2nd of May 1816 in London, where his father carried on business as a gun-maker. He had some schooling at Bexley, and was not at first intended for the artistic profession; but, developing a faculty in this line, he entered in 1834 the drawing class of Mr Sass, and in 1836 the school of the Royal Academy. His first exhibited picture appeared in 1837 at the Suffolk Street gallery. In 1838 he began exhibiting in the Academy, his subject being a "Spanish Girl"; altogether he sent twenty-seven works to this institution. In 1848 he became an associate and in 1860 a full member of the Academy: he had considerable means, apart from his profession. In 1857 he took a leading part in selecting and arranging the modern paintings in the Art-Treasures Exhibition in Manchester. His constitution being naturally frail, he went in 1853, with Dickens and Wilkie Collins, to Italy for a short trip, and in 1863 he visited Algeria. Here he benefited so far as his chronic lung-disease was concerned; but exposure to a cold wind while out riding brought on an attack of asthma, from which he died on the 26th of March 1863 at Algiers, near which city his remains were buried.

Egg was a gifted and well-trained painter of genre, chiefly in the way of historical anecdote, or of compositions from the poets and novelists. Among his principal pictures may be named: 1843, the "Introduction of Sir Piercie Shafton and Halbert Glendinning" (from Scott's _Monastery_); 1846, "Buckingham Rebuffed"; 1848, "Queen Elizabeth discovers she is no longer young"; 1850, "Peter the Great sees Catharine for the first time"; 1854, "Charles I. raising the Standard at Nottingham" (a study); 1855, the "Life and Death of Buckingham"; 1857 and 1858, two subjects from Thackeray's _Esmond_; 1858, "Past and Present, a triple picture of a faithless wife"; 1859, the "Night before Naseby"; 1860, his last exhibited work, the Dinner Scene from _The Taming of the Shrew_. The Tate Gallery contains one of his earlier pictures, Patricio entertaining two Ladies, from the _Diable boiteux_; it was painted in 1844.

Egg was rather below the middle height, with dark hair and a handsome well-formed face; the head of Peter the Great (in the picture of Peter and Catharine, which may be regarded as his best work, along with the Life and Death of Buckingham) was studied, but of course considerably modified, from his own countenance. He was manly, kind-hearted, pleasant, and very genial and serviceable among brother-artists; social and companionable, but holding mainly aloof from fashionable circles. As an actor he had uncommon talent. He appeared among Dickens's company of amateurs in 1852 in Lord Lytton's comedy _Not so Bad as we Seem_, and afterwards in Wilkie Collins's _Frozen Deep_, playing the humorous part of Job Want.

EGG (O.E. _aeg_, cf. Ger. _Ei_, Swed. _aegg_, and prob. Gr. [Greek: oon], Lat. _ovum_), the female reproductive cell or ovum of animals, which gives rise generally only after fertilization to the young. The largest eggs are those of birds; and this because, to the minute essential portion of the egg, or germ, from which the young bird grows, there is added a large store of food-material--the yolk and white of the egg--destined to nourish the growing embryo while the whole is enclosed within a hard shell.

The relative sizes of eggs depend entirely on the amount of the food-yolk thus enclosed with the germ; while the form and texture of the outer envelope are determined by the nature of the environment to which the egg is exposed. Where the food material is infinitesimal in quantity the egg is either not extruded--the embryo being nourished by the maternal tissues,--or it passes out of the parental body and gives rise at once to a free-living organism or "larva" (see LARVAL FORMS), as in the case of many lowly freshwater and marine animals. In such cases no "egg" in the usual sense of the term is produced.

The number of eggs periodically produced by any given individual depends on the risks of destruction to which they, and the young to which they give rise, are exposed: not more than a single egg being annually laid by some species, while with others the number may amount to millions.

_Birds' Eggs._--The egg of the bird affords, for general purposes, the readiest example of the modifications imposed on eggs by the external environment. Since it must be incubated by the warmth of the parent's body, the outer envelope has taken the form of a hard shell for the protection of the growing chick from pressure, while the dyes which commonly colour the surface of this shell serve as a screen to hide it from egg-eating animals.

Carbonate of lime forms the principal constituent of this shell; but in addition phosphate of lime and magnesia are also present. In section, this shell will be found to be made up of three more or less distinct crystalline layers, traversed by vertical canals, whereby the shell is made porous so as to admit air to the developing chick.

The outermost, or third, layer of this shell often takes the form of a glaze, as of porcelain, as for example in the burnished egg of the ostrich: or it may assume the character of a thick, chalky layer as in some cuckoos (_Guira_, _Crotophaga ani_), cormorants, grebes and flamingoes: while in some birds as in the auks, gulls and tinamous, this outer layer is wanting; yet the tinamous have the most highly glazed eggs of all birds, the second layer of the shell developing a surface even more perfectly burnished than that formed by the outermost, third layer in the ostrich.

While the eggs of some birds have the shell so thin as to be translucent, e.g. kingfisher, others display considerable thickness, the maximum being reached in the egg of the extinct _Aepyornis_.

Though in shape differing but little from that of the familiar hen's egg, certain well-marked modifications of form are yet to be met with. Thus the eggs of the plover are pear-shaped, of the sand-grouse more or less cylindrical, of the owls and titmice spherical and of the grebes biconical.

In the matter of coloration the eggs of birds present a remarkable range. The pigments to which this coloration is due have been shown, by means of their absorption spectra (Sorby, _Proc. Zool. Soc._, 1875), to be seven in number. The first of these, oorhodeine, is brown-red in tone, and rarely absent: the second and third, oocyanin, and banded oocyanin, are of a beautiful blue, and though differing spectroscopically give rise to the same product when oxidized: the fourth and fifth are yellow, and rufous ooxanthine, the former combining with oocyanin gives rise to the wonderful malachite green of the emu's egg, while the latter occurs only in the eggs of tinamous: the sixth is lichenoxanthine, a pigment not yet thoroughly known but present in the shells of all eggs having a peculiar brick-red colour. Still less is known of the seventh pigment which is, as yet, nameless. It is a substance giving a banded absorption spectrum, and which, mixed with other pigments, imparts an abnormally browner tint. The origin of these pigments is yet uncertain, but it is probable that they are derived from the haemoglobin or red colouring matter of the blood. This being so, then the pigments of the egg-shell differ entirely in their nature from those which colour the yolk or the feathers.

While many eggs are either colourless or of one uniform tint, the majority have the surface broken up by spots or lines, or a combination of both, of varying tints: the pigment being deposited as the egg passes down the lower portion of the oviduct. That the egg during this passage turns slowly on its long axis is shown by the fact that the spots and lines have commonly a spiral direction; though some of the markings are made during periods of rest, as is shown by their sharp outlines, movement giving a blurred effect. Where the egg is pyriform, the large end makes way for the smaller. Many eggs display, in addition to the strongly marked spots, more or fewer fainter spots embedded in a deeper layer of the shell, and hence such eggs are said to be "double-spotted," e.g. rails and plovers.

Among some species, as in birds of prey, the intensity of this coloration is said to increase with age up to a certain point, when it as gradually decreases. Frequently, especially where but two eggs are laid (Newton), all the dye will be deposited, sometimes on the first, sometimes on the last laid, leaving the other colourless. But although of a number of eggs in a "clutch"--as the full complement of eggs in a nest is called--no two are exactly alike, they commonly bear a very close resemblance. Among certain species, however, which lay several eggs, one of the number invariably differs markedly from the rest, as for example in the eggs of the house-sparrow or in those of the sparrow-hawk, where, of a clutch of six, two generally differ conspicuously from the rest. Differing though these eggs do from the rest of the clutch, all yet present the characters common to the species. But the eggs of some birds, such as the Australian swamp quail, _Synoecus australis_, present a remarkably wide range of variation in the matter of coloration, no two clutches being alike, the extremes ranging from pure white to eggs having a greenish ground colour and rufous spots or blotches. But a still more interesting illustration of variation equally marked is furnished by the chikor partridge (_Caccabis chukar_), since here the variation appears to be correlated with the geographical distribution of the species. Thus eggs taken in Greece are for the most part cream-coloured and unspotted; those from the Grecian Archipelago are generally spotted and blotched; while more to the eastward spots are invariably present, and the blotches attain their maximum development.

But in variability the eggs of the guillemot (_Lomvia troile_) exceed all others: both in the hue of the ground colour and in the form of the superimposed markings, these eggs exhibit a wonderful range for which no adequate explanation has yet been given.

Individual peculiarities of coloration are commonly reproduced, not only with this species but also in others, year after year.

Significance of colour.

The coloration of the egg bears no sort of relation to the coloration of the bird which lays it; but it bears on the other hand a more or less direct relation to the nature of the environment during incubation.

White eggs may generally be regarded as representing the primitive type of egg, since they agree in this particular with the eggs of reptiles. And it will generally be found that eggs of this hue are deposited in holes or in domed nests. So long indeed as nesting-places of this kind are used will the eggs be white. And this because coloured eggs would be invisible in dimly lighted chambers of this description, and therefore constantly exposed to the risk of being broken by the sitting bird, or rolling out of reach where the chamber was large enough to admit of this, whereas white eggs are visible so long as they can be reached by the faintest rays of light. Pigeons invariably lay white eggs; and while some deposit them in holes others build an open nest, a mere platform of sticks. These exceptions to the rule show that the depredations of egg-eating animals are sufficiently guarded against by the overhanging foliage, as well as by the great distance from the ground at which the nest is built. Birds which have reverted to the more ancient custom of nesting in holes after having developed pigmented eggs, have adopted the device of covering the shell with a layer of chalky matter (e.g. puffins), or, to put the case more correctly, they have been enabled to maintain survival after their return to the more ancient mode of nidification, because this reversion was accompanied by the tendency to cover the pigmented surface of the shell with this light-reflecting chalky incrustation.

Eggs which are deposited on the bare ground, or in other exposed situations, are usually protectively coloured: that is to say, the hue of the shell more or less completely harmonizes with the ground on which the egg is placed. The eggs of the plover tribe afford the most striking examples of this fact.

But the majority of birds deposit their eggs in a more or less elaborately constructed nest, and in such cases the egg, so far from being protectively coloured, often displays tints that would appear calculated rather to attract the attention of egg-stealing animals; bright blue or blue spotted with black being commonly met with. It may be, however, that coloration of this kind is less conspicuous than is generally supposed, but in any case the safety of the egg depends not so much on its coloration as on the character of the nest, which, where protective devices are necessary, must harmonize sufficiently with its surroundings to escape observation from prowling egg-stealers of all kinds.

The size of the egg depends partly on the number produced and partly on the conditions determining the state of the young bird at hatching: hence there is a great disparity in the relative sizes of the eggs of different birds. Thus it will be found that young birds which emerge in the world blind, naked and helpless are the product of relatively small eggs, while on the contrary young hatched from relatively large eggs are down-clad and active from birth.

The fact that the eggs must be brooded by the parent is also a controlling factor in so far as number is concerned, for no more can be hatched than can be covered by the sitting bird. Other factors, however, less understood, also exercise a controlling influence in this matter. Thus the ostrich lays from 12 to 16, the teal 15, the partridge 12-20, while among many other species the number is strictly limited, as in the case of the hornbills and guillemots, which lay but a single egg; the apteryx, divers, petrels and pigeons never lay more than 2, while the gulls and plovers never exceed 4. Tropical species are said to lay fewer eggs than their representatives in temperate regions, and further immature birds lay more and smaller eggs than when fully adult.

Partly owing to the uniformity of shape, size and texture of the shell, the eggs of birds are by no means easy to distinguish, except in so far as their family resemblances are concerned: that is to say, except in particular cases, they cannot be specifically distinguished, and hence they are of but little or no value for the purposes of classification.

Save only among the megapodes, all birds brood their eggs, the period of incubation varying from 13 days, as in small passerine birds, to 8 weeks, as in the cassowary, though eggs of the rhea and of _Struthio_ hatch in from 5 to 6 weeks. But the megapodes deposit their eggs in mounds of decaying vegetable matter or in sand in the neighbourhood of hot springs, and there without further apparent care leave them. Where the nestling is active from the moment of hatching the eggs have a relatively longer incubation period than in cases where the nestlings are for a long while helpless.

_Eggs of Mammals._--Only in the spiny ant-eater, or _Echidna_, and the duck-billed platypus, or _Ornithorhynchus_, among the Mammalia, are the eggs provided with a large store of yolk, enclosed within a shell, and extruded to undergo development apart from the maternal tissues. In the case of the echidna the eggs, two in number, are about as large as those of a sparrow, similar in shape, and have a white, parchment-like shell. After expulsion they are transferred by the beak of the mother to a pouch resembling that of the marsupial kangaroos, and there they undergo development. The _Ornithorhynchus_, on the other hand, lays from two to four eggs, which in size and general appearance resemble those of the echidna. They are, however, deposited in a loosely constructed nest at the end of a long burrow and there brooded. In Marsupials, the eggs are smaller than those of _Echidna_ and _Ornithorhynchus_, and they contain a larger proportion of yolk than occurs in higher mammals.

_Eggs of Reptiles._--The eggs of reptiles are invariably provided with a large amount of food yolk and enclosed with a firm test or shell, which though generally parchment-like in texture may be calcareous as in birds, as, for example, in many of the tortoises and turtles and in the crocodiles.

Among reptiles the egg is always white or yellowish, while the number laid often far exceeds that in the case of birds. The tuatara of New Zealand, however, lays but ten--white hard-shelled, long and oval--at intervals between November and January. The long intervals between the appearance of the successive eggs is a characteristic feature of the reptiles, but is met with among the birds only in the megapodes, which, like the reptiles, do not "brood" their eggs.

Among the Chelonia the number of eggs varies from two to four in some of the tortoises, to 200 in some of the turtles: while in the crocodiles between 20 and 30 are produced, hard-shelled and white.

The eggs of the lizards are always white or yellowish, and generally soft-shelled; but the geckos and the green lizard lay hard-shelled eggs. Many of the soft-shelled eggs are remarkable for the fact that they increase in size after extrusion, owing to the stretching of the membranous shell by the growing embryo. In the matter of number lizards are less prolific than many of the Chelonia, a dozen eggs being the general number, though as many as thirty may be produced at a time, as in the case of the common chameleon.

While as a general rule the eggs of lizards are laid in burrows or buried, some are retained within the body of the parent until the young are ready to emerge; or they may even hatch within the oviduct. This occurs with some chameleons and some lizards, e.g. the slow-worm. The common English lizard is also viviparous. Normally the young leaves the egg immediately after its extrusion, but if by any chance this extrusion is delayed they escape while yet in the oviduct.

The majority of the snakes lay eggs, but most of the vipers and the aquatic snakes are viviparous, as also are a few terrestrial species. The shell of the egg is always soft and parchment-like. As a rule the number of eggs produced among the snakes is not large, twenty or thirty being common, but some species of python lay as many as a hundred. Generally, among the oviparous snakes the eggs are buried, but some species of boas jealously guard them, enclosing them within the coils of the body.

_Eggs of Amphibia._--Among the amphibia a greater variety obtains in the matter of the investment of the egg, as well as in the number, size and method of their disposal. The outer covering is formed by a toughening of the surface of a thick gelatinous coat which surrounds the essential parts of the egg. This coat in many species of salamander--using this name in the wide sense--is produced into threads which serve either to anchor the eggs singly or to bind them together in bunches.

Viviparity occurs both among the limbless and the tailed Amphibia, the eggs hatching before they leave the oviduct or immediately after extrusion. The number of young so produced is generally not large, but the common salamander (_Salamandra maculosa_) may produce as many as fifty at a birth, though fifteen is the more normal figure. When the higher number is reached the young are relatively small and weak.

As a rule among the Amphibia the young leave the egg in the form of larvae, generally known as "tadpoles"; but many species produce eggs containing a sufficient amount of food material to enable the whole of the larval phase to be completed before hatching.

Among the tailless Amphibia (frogs and toads) there are wide differences in the number of eggs produced, while the methods by which these eggs are disposed of present a marvellous variety.

As a rule vast quantities of eggs are shed by the female into the water in the form of "spawn." In the common toad as many as 7000 eggs may be extruded at a time. These leave the body in the form of two long strings--one from each oviduct--of translucent globules, gelatinous in texture, and enclosing a central sphere of yolk, the upper pole of which is black. The spawn of the common frog differs from that of the toad in that the eggs all adhere to form a huge jelly-like mass. But in many species the number of eggs produced are few; and these may be sufficiently stored with food-yolk to allow of the tadpole stage being passed before hatching, as in frogs of the genus _Hylodes_. In many cases the eggs are deposited out of the water and often in quite remarkable ways.

_Eggs of Fishes._--The eggs of fishes present an extremely wide range of form, and a no less extensive range in the matter of number. Both among the cartilaginous and bony fishes viviparity occurs. Most of the sharks and rays are viviparous, but in the oviparous species the eggs present some interesting and peculiar forms. Large in size, the outer coat or "shell" is in all cases horn-like and flexible, but differs greatly in shape. Thus in the egg of the larger spotted dog-fish it is oblong in shape, flattened from side to side, and has the angles produced into long, slender tendrils. As the egg is laid the lower tendrils project from the vent, and the mother rubs herself against some fixed body. The tendrils soon catch fast in some slight projection, when the egg is dragged forth there to remain till hatching takes place. A couple of narrow slits at each corner of the upper end serve to admit fresh water to the imprisoned embryo during the later stages of development; when development is complete escape is made through the end of the shell. In the rays or "skates," long spines take the place of tendrils, the egg simply resting at the bottom of the sea. The empty egg-cases of the rays are often found on the seashore, and are known as "Mermaids' purses." The egg of the Port Jackson shark (_Cestracion_) is of enormous size, pear-shaped, and provided with a spiral flange extending along the whole length of the capsule. In the _Chimaera_ the egg is long, more or less spindle-shaped, and produced on each side into a broad flange having a fringed edge, so that the whole bears a close resemblance to a long leaf, broad and notched at one end, pointed at the other. This likeness to the seaweed among which it rests is doubtless a protective device, akin to that of protectively coloured birds' eggs.

Among the bony fishes the eggs generally take the form of small spheres, enclosed within a tough membrane or capsule. But they present many important differences, being in some fishes heavy and remaining at the bottom of the water, in other light and floating on the surface. While in some species they are distributed separately, in others they adhere together in masses. The eggs of the salmon, for example, are heavy, hard and smooth, and deposited separately in a trough dug by the parent and afterwards covered to prevent them from being carried away by the stream. In the perch they are adhesive and form long band-like masses of spawn adhering to water-plants. In the gobies the egg is spindle-shaped, and attached by one end by means of a network of fibres, resembling rootlets; while in the smelt the egg is loosely suspended by a membrane formed by the peeling off of a part of the outer sheath of the capsule. The eggs of the garfish (_Belone vulgaris_) and of the flying-fish of the genus _Exocoetus_, attach themselves to foreign objects, or to one another, by means of threads or cords developed at opposite poles of the egg.

Among a number of fishes the eggs float at the surface of the sea, often in enormous masses, when they are carried about at the mercy of tides and currents. An idea of the size which such masses attain may be gathered from the fact that the spawn of the angler-fish, _Lophius piscatorius_, takes the form of a sheet from 2 to 3 ft. wide, and 30 ft. long. Another remarkable feature of these floating eggs is their transparency, inasmuch as they are extremely difficult to see, and hence they probably escape the rapacious maws of spawn-eating animals. The cod tribe and flat-fishes lay floating eggs of this description.

The maximum number of eggs laid by fishes varies greatly, some species laying relatively few, others an enormous number. But in all cases the number increases with the weight and age of the fish. Thus it has been calculated that the number laid by the salmon is roughly about 1000 to every pound weight of the fish, a 15 lb. salmon laying 15,000 eggs. The sturgeon lays about 7,000,000; the herring 50,000; the turbot 14,311,000; the sole 134,000; the perch 280,000. Briefly, the number is greatest where the risks of destruction are greatest.

The eggs of the degenerate fishes known as the lampreys and hag-fishes are remarkable for the fact that in the latter they are large in size, cylindrical in shape, and provided at each end with hooklets whereby they adhere one to another; while in the lampreys they are extremely small and embedded in a jelly.

_Molluscs._--Among the Mollusca, Crustacea and Insecta yolk-stored eggs of very remarkable forms are commonly produced.

In variety, in this connexion, the Mollusca must perhaps be given the first place. This diversity, indeed, is strikingly illustrated by the eggs of the Cephalopoda. In the squids (_Loligo_), for example, the eggs are enclosed in long cylindrical cases, of which there are several hundreds, attached by one end to a common centre; the whole series looking strangely like a rough mop-head. Each case, in such a cluster, contains about 250 eggs, or about 40,000 in all. By way of contrast the eggs of the true cuttle-fish (_Sepia_) are deposited separately, each enclosed in a tough, black, pear-shaped capsule which is fastened by a stalk to fronds of sea-weed or other object. They appear to be extruded at short intervals, till the full complement is laid, the whole forming a cluster looking like a bunch of grapes. The octopus differs yet again in this matter, its eggs being very small, berry-like, and attached to a stalk which runs through the centre of the mass.

The eggs of the univalve Mollusca are hardly less varied in the shapes they take. In the common British _Purpura lapillus_ they resemble delicate pink grains of rice set on stalks; in _Busycon_ they are disk-shaped, and attached to a band nearly 3 ft. long. The eggs of the shell-bearing slugs (_Testacella_) are large, and have the outer coat so elastic that if dropped on a stone floor they will rebound several inches; while some of the snails (_Bulimus_) lay eggs having a white calcareous and slightly iridescent shell, in size and shape closely resembling the egg of the pigeon. Some are even larger than the egg of the wood-pigeon. The beautiful violet-snail (_Ianthina_)--a marine species--carries its eggs on the under side of a gelatinous raft. No less remarkable are the eggs of the whelk; since, like those of the squids, they are not laid separately but enveloped in capsules, and these to the number of many hundreds form the large, ball-like masses so commonly met with on the seashore. When the eggs in these capsules hatch, the crowd of embryos proceed to establish an internecine warfare, devouring one another till only the strongest survives!

With the Mollusca, as with other groups of animals, where the eggs are exposed to great risks they are small, produced in great numbers, and give rise to larvae. This is well illustrated by the common oyster which annually disperses about 60,000,000 eggs. But where the risk of destruction is slight, the eggs are large and produce young differing from the parent only in size, as in the case of the pigeon-like eggs of _Bulimus_.

_Crustaceans._--Among the higher Crustacea, as a rule, the eggs are carried by the female, attached to special appendages on the under side of the body. But in some--Squillas--they are deposited in burrows. Generally they are relatively small so that the young which emerge therefrom differ markedly in appearance from the parents, but in deep-sea and freshwater species the eggs are large, when the young, on emerging, differ but little from the adults in appearance.

_Insects, &c._--The eggs of insects though minute, are also remarkable for the great variety of form which they present, while they are frequently objects of great beauty owing to the sculptured markings of the shell. They are generally laid in clusters, either on the ground, on the leaves of plants, or in the water. Some of the gnats (_Culex_) lay them on the water. Cylindrical in shape they are packed closely together, set on end, the whole mass forming a kind of floating raft. Frequently, as in the case of the stick and leaf insect, the eggs are enclosed in capsules of very elaborate shapes and highly ornamented.

As to the rest of the Invertebrata--above the Protozoa the eggs are laid in water, or in damp places. In the former case they are as a rule small, and give rise to larvae; while eggs hatched on land are sometimes enclosed in capsules, "cocoons," as in the case of the earthworm, where this capsule is filled with a milky white fluid, of a highly nutritious character, on which the embryos feed.

Among some invertebrates two different kinds of eggs are laid by the same individual. The water-flea, _Daphnia_ (a crustacean), lays two kinds of eggs known as "summer" and "winter" eggs. The summer eggs are carried by the female in a "brood-pouch" on the back. The "winter" eggs, produced at the approach of winter, differ markedly in appearance from the summer eggs, being larger, darker in colour, thicker shelled, and enclosed in a capsule formed from the shell or carapace, of the parent's body. "Winter eggs," however, may be produced in the height of summer. While the "summer eggs" are unfertilized, the winter eggs are fertilized by the male, and possess the remarkable power of lying dormant for months or even years before they develop. The production of these two kinds of eggs is a device to overcome the cold of winter, or the drying up of the pools in which the species lives, during the heat of the summer. The power of resistance which such eggs possess may be seen in the fact that a sample of mud which had been kept dry for ten years still contained living eggs. In deep water where neither drought nor winter cold can seriously affect the _Daphnias_, they propagate all the year round by unfertilized "summer" eggs.

BIBLIOGRAPHY.--For further details on this subject the following
authors should be consulted:--_Mammals_: F. E. Beddard, "Remarks on
the Ovary of Echidna," _Proc. Roy. Phys. Soc. Edin._ vol. viii.
(1885); W. H. Caldwell, "The Embryology of Monotremata and
Marsupialia," _Phil. Trans. Roy. Soc._ vol. 178 (1887); E. B. Poulton,
"The Structures connected with the Ovarian Ovum of the Marsupialia and
Monotremata," _Quart. Journ. Micros. Sci._ vol. xxiv. (1884). _Birds,
Systematic_:--H. Seebohm, _Coloured Figures of the Eggs of British
Birds_ (1896); A. Newton, _Ootheca Wooleyana_ (1907); E. Oates, _Cat.
Birds' Eggs Brit. Mus._ (appearing), vols. i.-iv. published.
_General_:--A. Newton, _Dictionary of Birds_ (1896). _Colouring
matter_:--Newbegin, _Colour in Nature_ (1898). _Reptiles and
Amphibia_:--H. Gadow, "Reptiles," _Camb. Nat. Hist._ (1901); G. A.
Boulenger, "The Tailless Batrachians of Europe," _Ray Soc._ (1896).
_Fishes_:--Bridge and Boulenger, "Fishes, Ascidians, &c.," _Camb. Nat.
Hist._ (1904); B. Dean, _Fishes Living and Fossil_ (1895); J. T.
Cunningham, _Marketable Marine Fishes_ (1896). _Invertebrate_:--G. H.
Carpenter, _Insects. Their Structure and Life_ (1899); L. C. Miall, _A
History of Aquatic Insects_ (1895); T. R. R. Stebbing, _Crustacea_,
Internat. Sci. series (1893); M. C. Cooke, "Mollusca," _Camb. Nat.
Hist._ (1906). For further references to the above and other
Invertebrate groups see various text-books on Entomology, Zoology.
(W. P. P.)

EGGENBERG, HANS ULRICH VON, PRINCE (1568-1634), Austrian statesman, was a son of Siegfried von Eggenberg (d. 1594), and began life as a soldier in the Spanish service, becoming about 1596 a trusted servant of the archduke of Styria, afterwards the emperor Ferdinand II. Having become a Roman Catholic, he was soon the chancellor and chief adviser of Ferdinand, whose election as emperor he helped to secure in 1619. He directed the imperial policy during the earlier part of the Thirty Years' War, and was in general a friend and supporter of Wallenstein, and an opponent of Maximilian I., duke of Bavaria, and of Spain. He was largely responsible for Wallenstein's return to the imperial service early in 1632, and retired from public life just after the general's murder in February 1634, dying at Laibach, on the 18th of October 1634. Eggenberg's influence with Ferdinand was so marked that it was commonly said that Austria rested upon three hills (_Berge_): Eggenberg, Questenberg and Werdenberg. He was richly rewarded for his services to the emperor. Having received many valuable estates in Bohemia and elsewhere, he was made a prince of the Empire in 1623, and duke of Krumau in 1625.

See H. von Zwiedineck-Sudenhorst, _Hans Ulrich, Furst von Eggenberg_
(Vienna, 1880); and F. Mares, _Beitrage zur Geschichte der Beziehungen
des Fursten J. U. von Eggenberg zu Kaiser Ferdinand II und zu
Waldstein_ (Prague, 1893).

EGGER, EMILE (1813-1885), French scholar, was born in Paris on the 18th of July 1813. From 1840 till 1855 he was assistant professor, and from 1855 till his death professor of Greek literature in the Faculte; des Lettres at Paris University. In 1854 he was elected a member of the Academie des Inscriptions and in 1873 of the Conseil superieur de l'instruction publique. He was a voluminous writer, a sound and discerning scholar, and his influence was largely responsible for the revival of the study of classical philology in France. His most important works were _Essai sur l'histoire de la critique chez les Grecs_ (1849), _Notions elementaires de grammaire comparee_ (1852), _Apollonius Dyscole, essai sur l'histoire des theories grammalicales dans l'antiquite_ (1854), _Memoires de litterature ancienne_ (1862), _Memoires d'histoire ancienne et de philologie_ (1863), _Les Papyrus grecs du Musee du Louvre et de la Bibliotheque Imperiale_ (1865), _Etudes sur les traites publics chez les Grecs et les Romains_ (1866), _L'Hellenisme en France_ (1869), _La Litterature grecque_ (1890). He was also the author of _Observations et reflexions sur le developpement de l'intelligence et du langage chez les enfants_ (1879). Egger died in Paris on the 1st of September 1885.

EGGLESTON, EDWARD (1837-1902), American novelist and historian, was born in Vevay, Indiana, on the 10th of December 1837, of Virginia stock. Delicate health, by which he was more or less handicapped throughout his life, prevented his going to college, but he was naturally a diligent student. He was a Methodist circuit rider and pastor in Indiana and Minnesota (1857-1866); associate editor (1866-1867) of _The Little Corporal_, Chicago; editor of _The National Sunday School Teacher_, Chicago (1867-1870); literary editor and later editor-in-chief of _The Independent_, New York (1870-1871); and editor of _Hearth and Home_ in 1871-1872. He was pastor of the church of Christian Endeavour, Brooklyn, in 1874-1879. From 1880 until his death on the 2nd of September 1902, at his home on Lake George, New York, he devoted himself to literary work. His fiction includes _Mr Blake's Walking Stick_ (1869), for children; _The Hoosier Schoolmaster_ (1871); _The End of the World_ (1872); _The Mystery of Metropolisville_ (1873); _The Circuit Rider_ (1874); _Roxy_ (1878); The _Hoosier Schoolboy_ (1883); _The Book of Queer Stories_ (1884), for children; _The Graysons_ (1888), an excellent novel; _The Faith Doctor_ (1891); and _Duffels_ (1893), short stories. Most of his stories portray the pioneer manners and dialect of the Central West, and the _Hoosier Schoolmaster_ was one of the first examples of American local realistic fiction; it was very popular, and was translated into French, German and Danish. During the last third of his life Eggleston laboured on a _History of Life in the United States_, but he lived to finish only two volumes--_The Beginners of a Nation_ (1896) and _The Transit of Civilization_ (1900). In addition he wrote several popular compendiums of American history for schools and homes.

See G. C. Eggleston, _The First of the Hoosiers_ (Philadelphia, 1903),
and Meredith Nicholson, _The Hoosiers_ (1900).

His brother GEORGE CARY EGGLESTON (1839- ), American journalist and author, served in the Confederate army; was managing editor and later editor-in-chief of _Hearth and Home_ (1871-1874); was literary editor of the _New York Evening Post_ (1875-1881), literary editor and afterwards editor-in-chief of the New York _Commercial Advertiser_ (1884-1889), and editorial writer for _The World_ (New York) from 1889 to 1900. Most of his books are stories for boys; others, and his best, are romances dealing with life in the South especially in the Virginias and the Carolinas--before and during the Civil War. Among his publications may be mentioned: _A Rebel's Recollections_ (1874); _The Last of the Flatboats_ (1900); _Camp Venture_ (1900); _A Carolina Cavalier_ (1901); _Dorothy South_ (1902); _The Master of Warlock_ (1903); _Evelyn Byrd_ (1904); _A Daughter of the South_ (1905); _Blind Alleys_ (1906); _Love is the Sum of it all_ (1907); _History of the Confederate War_ (1910); and _Recollections of a Varied Life_ (1910).

EGHAM, a town in the Chertsey parliamentary division of Surrey, England, on the Thames, 21 m. W.S.W. of London by the London & South Western railway. Pop. (1901) 11,895. The church of St John the Baptist is a reconstruction of 1817; it contains monuments by John Flaxman. Above the right bank of the river a low elevation, Cooper's Hill, commands fine views over the valley, and over Windsor Great Park to the west. On the hill was the Royal Indian Civil Engineering College, commonly called Cooper's Hill College, of which Sir George Tomkyns Chesney was the originator and first president (1871). It educated men for the public works, accounts, railways and telegraph departments of India, and included a school of forestry; but it was decided, in the face of some opposition, to close it in 1906, on the theory that it was unnecessary for a college with such a specialized object to be maintained by the government, in view of the readiness with which servants for these departments could be recruited elsewhere. Part of the organization, including the school of forestry, was transferred to Oxford University. Cooper's Hill gives name to a famous poem of Sir John Denham (1642). A large and handsome building houses the Royal Holloway College for Women (1886), founded by Thomas Holloway; in the neighbourhood is the sanatorium of the same founder (1885) for the treatment of mental ailments, accommodating about 250 patients. The college for women, surrounded by extensive grounds, commands a wide view from the wooded slope on which it stands. The recreation hall, with its fine art collection, is the most notable room in this handsome building, which can receive 250 students. Within the parish, bordering the river, is the field of Runnymede, which, with Magna Charta Island lying off it, is famous in connexion with the signature of the charter by King John. Virginia Water, a large and picturesque artificial lake to the south of Windsor Great Park, is much frequented by visitors. It was formed under the direction of the duke of Cumberland, about 1750, and was the work of the brothers Thomas and Paul Sandby.

EGIN (Armenian _Agn_, "the spring"), an important town in the Mamuret el-Aziz vilayet of Asiatic Turkey (altitude 3300 ft.). Pop. about 20,000, fairly equally divided between Armenian Christians and Moslems. It is picturesquely situated in a theatre of lofty, abrupt rocks, on the right bank of the western Euphrates, which is crossed by a wooden bridge. The stone houses stand in terraced gardens and orchards, and the streets are mere rock ladders. Egin was settled by Armenians who emigrated from Van in the 11th century with Senekherim. On the 8th of November 1895 and in the summer of 1896 many Armenians were massacred here. (D. G. H.)

EGLANTINE (E. Frisian, _egeltiere_; Fr. _aiglantier_), a plant-name of which Dr R. C. A. Prior (_Popular Names of British Plants_, p. 70) says that it "has been the subject of much discussion, both as to its exact meaning and as to the shrub to which it properly belongs." The eglantine of the herbalists was the sweet-brier, _Rosa rubiginosa_. The signification of the word seems to be thorn-tree or thorn-bush, the first two syllables probably representing the Anglo-Saxon _egla_, _egle_, a prick or thorn, while the termination is the Dutch _tere_, _taere_, a tree. Eglantine is frequently alluded to in the writings of English poets, from Chaucer downwards. Milton, in _L'Allegro_, is thought by the term "twisted eglantine" to denote the honeysuckle, _Lonicera Periclymenum_, which is still known as eglantine in north-east Yorkshire.

EGLINTON, EARLS OF. The title of earl of Eglinton has been held by the famous Scottish family of Montgomerie since 1508. The attempts made to trace the descent of this house to Roger of Montgomery, earl of Shrewsbury (d. 1094), one of William the Conqueror's followers, will not bear examination, and the sure pedigree of the family only begins with Sir John Montgomerie, lord of Eaglesham, who fought at the battle of Otterbourne in 1388 and died about 1398. His grandson, Sir Alexander Montgomerie (d. c. 1460), was made a lord of the Scottish parliament about 1445 as Lord Montgomerie, and Sir Alexander's great-grandson Hugh, the 3rd lord (c. 1460-1545), was created earl of Eglinton, or Eglintoun, in 1508. Hugh, who was a person of importance during the minority of James V., was succeeded by his grandson Hugh (d. 1546), and then by the latter's son Hugh (c. 1531-1585), who became 3rd earl of Eglinton. This nobleman was a firm supporter of Mary queen of Scots, for whom he fought at Langside, and of the Roman Catholic Church; his son and successor, Hugh, was murdered in April 1586 by the Cunninghams, a family with which his own had an hereditary blood feud. In 1612, by the death of Hugh, the 5th earl, the male line of the Montgomeries became extinct.

Having no children Earl Hugh had settled his title and estates on his cousin, Sir Alexander Seton of Foulstruther (1588-1661), a younger son of Robert Seton, 1st earl of Wintoun (c. 1550-1603), and his wife Margaret, daughter of the 3rd earl of Eglinton. Alexander, who thus became the 6th earl of Eglinton and took the name of Montgomerie, was commonly called Greysteel; he was a prominent Covenanter and fought against Charles I. at Marston Moor. Later, however, he supported the cause of Charles II., and fell into the hands of Cromwell, who imprisoned him. His fifth son, Robert Montgomerie (d. 1684), a soldier of distinction, fought against Cromwell at Dunbar and at Worcester, afterwards escaping from the Tower of London and serving in Denmark. Robert's elder brother, Hugh, 7th earl of Eglinton (1613-1669), who also fought against Cromwell, was the grandfather of Alexander, the 9th earl (c. 1660-1729), who married, for his third wife, Susannah (1689-1780), daughter of Sir Archibald Kennedy, Bart., of Culzean, a lady celebrated for her wit and beauty. Alexander, the 10th earl (1723-1769), a son of the 9th earl, was one of the first of the Scottish landowners to carry out improvements on his estates. He was shot near Ardrossan by an excise officer named Mungo Campbell on the 24th of October 1769. His brother and successor, Archibald, the 11th earl (1726-1796), raised a regiment of Highlanders with which he served in America during the Seven Years' War. As he left no male issue he was succeeded in the earldom by his kinsman Hugh Montgomerie (1739-1819), a descendant of the 6th earl, who was created a peer of the United Kingdom as Baron Ardrossan in 1806. Before succeeding to the earldom Hugh had served in the American war and had been a member of parliament; after this event he began to rebuild Eglinton castle on a magnificent scale and to construct a harbour at Ardrossan.

This earl's successor was his grandson, Archibald William, the 13th earl (1812-1861), who was born at Palermo on the 29th of September 1812. His father was Archibald, Lord Montgomerie (1773-1814), the eldest son of the 12th earl, and his mother was Mary (d. 1848), a daughter of the 11th earl. Educated at Eton, the young earl's main object of interest for some years was the turf; he kept a large racing stud and won success and reputation in the sporting world. In 1839 his name became more widely known in connexion with the famous tournament which took place at Eglinton castle and is said to have cost him L30,000 or L40,000. This was made the subject of much ridicule and was partly spoiled by the unfavourable weather, the rain falling in torrents. Yet it was a real tournament and the "knights" broke their spears in the orthodox way. Prince Louis Napoleon (Napoleon III.) took part in it, and Lady Seymour, a daughter of Thomas Sheridan and the wife of Lord Seymour, afterwards 12th duke of Somerset, was the queen of beauty. A list of the challengers with an account of the jousts and the melee will be found in the volume on the tournament written by John Richardson, with drawings by J. H. Nixon. It is also described by Disraeli in _Endymion_. Eglinton was a staunch Tory, and in February 1852 he became lord-lieutenant of Ireland under the earl of Derby. He retired with the ministry in the following December, having by his princely hospitality made himself one of the most popular of Irish viceroys. When Derby returned to office in February 1858 he was again appointed lord-lieutenant, and he discharged the duties of this post until June 1859. In this year he was created earl of Winton, an earldom which had been held by his kinsfolk, the Setons, from 1600 until 1716, when George Seton, the 5th earl (c. 1678-1749), was deprived of his honours for high treason. The earl died on the 4th of October 1861, and was succeeded by his eldest son Archibald William (1841-1892). When this earl died in 1892 his younger brother George Arnulph (b. 1848) became 15th earl of Eglinton and 3rd earl of Winton.

See Sir W. Fraser, _Memorials of the Montgomeries, earls of Eglinton_
(1859).

EGMONT, EARLS OF. John Perceval, 1st earl of Egmont (1683-1748), Irish politician, and partner with J. E. Oglethorpe in founding the American colony of Georgia, was created earl in 1733. He claimed descent from the Egmonts of Flanders, but his title was taken from the place in County Cork where the family residence stood. Its name of Burton House, and that of Burton manor which formed part of the family estates, were a reminiscence of Burton in Somerset, where was the earlier English family property of his great-great-grandfather Richard Perceval (1550-1620), Burghley's secret agent, and author of a Spanish dictionary published in 1591, whose son Sir Philip Perceval (1605-1647) acquired the Irish estates by judicious use of his opportunities as commissioner for land titles and of his interest at court. Sir Philip's son John, grandfather of the 1st earl, was made a baronet in 1661. The first earl of Egmont (who had been made Baron Perceval in 1715, and Viscount Perceval in 1723) is chiefly important for his connexion with the colonization of Georgia, and for his voluminous letters and writings on biography and genealogy.

John Perceval, 2nd earl of Egmont (1711-1770), his eldest son, was an active politician, first lord of the admiralty (1763-1766), and political pamphleteer, and like his father an ardent genealogist. He was twice married, and had eight sons and eight daughters. One of his younger sons was Spencer Perceval, prime minister of England. His eldest son succeeded as 3rd earl, and the eldest by his second marriage (with Catherine Compton, baroness of Arden in Ireland) was in 1802 created Baron Arden of the United Kingdom, a title which subsequently became merged in the Egmont earldom.

EGMONT (EGMOND), LAMORAL, COUNT OF, prince of Gavre (1522-1568), was born in Hainaut in 1522. He was the younger of the two sons of John IV., count of Egmont, by his wife Francoise of Luxemburg, princess of Gavre. On the death of his elder brother Charles, about 1541, he succeeded to his titles and estates. In this year he served his apprenticeship as a soldier in the expedition of the emperor Charles V. to Algiers, distinguishing himself in the command of a body of cavalry. In 1544 he married Sabina, sister of the elector palatine Frederick III., and the wedding was celebrated at Spires with great pomp in the presence of the emperor and his brother Ferdinand, afterwards emperor. Created knight of the Golden Fleece in 1546, he accompanied Philip of Spain in his tour through the Netherland towns, and in 1554 he went to England at the head of a special embassy to ask the hand of Mary of England for Philip, and was afterwards present at the wedding ceremony at Winchester. In the summer of 1557 Egmont was appointed commander of the Flemish cavalry in the war between Spain and France; and it was by his vehement persuasion that the battle of St Quentin was fought. The victory was determined by the brilliant charge that he led against the French. The reputation which he won at St Quentin was raised still higher in 1558, when he encountered the French army under de Thermes at Gravelines, on its march homewards after the invasion of Flanders, totally defeated it, and took Marshal de Thermes prisoner. The battle was fought against the advice of the duke of Alva, and the victory made Alva Egmont's enemy. But the count now became the idol of his countrymen, who looked upon him as the saviour of Flanders from the devastations of the French. He was nominated by Philip stadtholder of Flanders and Artois. At the conclusion of the war by the treaty of Cateau Cambresis, Egmont was one of the four hostages selected by the king of France as pledges for its execution.