Chapter XXXVIII

LAWS WHICH REGULATE THE GEOGRAPHICAL DISTRIBUTION OF SPECIES--_continued_.

Geographical distribution of animals--Buffon on specific
distinctness of quadrupeds of Old and New World--Doctrine of
"natural barriers"--Different regions of indigenous
mammalia--Europe--Africa--India, and Indian
Archipelago--Australia--North and South America--Quadrupeds in
islands--Range of the Cetacea--Dispersion of quadrupeds--Their
powers of swimming--Migratory instincts--Drifting of animals on
ice-floes--On floating islands of drift-timber--Migrations of
Cetacea--Habitations of birds--Their migrations and facilities of
diffusion--Distribution of reptiles, and their power of
dissemination.

_Geographical distribution of animals._--Although in speculating on "philosophical possibilities," said Buffon, "the same temperature might have been expected, all other circumstances being equal, to produce the same beings in different parts of the globe, both in the animal and vegetable kingdoms, yet it is an undoubted fact, that when America was discovered, its indigenous quadrupeds were all dissimilar to those previously known in the Old World. The elephant, the rhinoceros, the hippopotamus, the camelopard, the camel, the dromedary, the buffalo, the horse, the ass, the lion, the tiger, the apes, the baboons, and a number of other mammalia, were nowhere to be met with on the new continent; while in the old, the American species, of the same great class, were nowhere to be seen--the tapir, the lama, the pecari, the jaguar, the couguar, the agouti, the paca, the coati, and the sloth."

These phenomena, although few in number relatively to the whole animate creation, were so striking and so positive in their nature, that the great French naturalist caught sight at once of a general law in the geographical distribution of organic beings, namely, the limitation of groups of distinct species to regions separated from the rest of the globe by certain natural barriers. It was, therefore, in a truly philosophical spirit that, relying on the clearness of the evidence obtained respecting the larger quadrupeds, he ventured to call in question the identifications announced by some contemporary naturalists of species of animals said to be common to the southern extremities of America and Africa.[871]

The migration of quadrupeds from one part of the globe to another, observes Dr. Prichard, is prevented by uncongenial climates and the branches of the ocean which intersect continents. "Hence, by a reference to the geographical site of countries, we may divide the earth into a certain number of regions fitted to become the abodes of particular groups of animals, and we shall find, on inquiry, that each of these provinces, thus conjecturally marked out, is actually inhabited by a distinct nation of quadrupeds."[872] It will be observed that the language of Buffon respecting "natural barriers," which has since been so popular, would be wholly without meaning if the geographical distribution of organic beings had not led naturalists to adopt very generally _the doctrine of specific centres_, or, in other words, to believe that each species, whether of plant or animal, originated in a single birth-place. Reject this view, and the fact that not a single native quadruped is common to Australia, the Cape of Good Hope, and South America, can in no ways be explained by adverting to the wide extent of intervening ocean, or to the sterile deserts, or the great heat or cold of the climates, through which each species must have passed, before it could migrate from one of those distant regions to another. It might fairly be asked of one who talked of impassable barriers, why the same kangaroos, rhinoceroses, or lamas, should not have been created simultaneously in Australia, Africa, and South America! The horse, the ox, and the dog, although foreign to these countries until introduced by man, are now able to support themselves there in a wild state, and we can scarcely doubt that many of the quadrupeds at present peculiar to Australia, Africa, and South America, might have continued in like manner to inhabit each of the three continents had they been indigenous or could they once have got a footing there as new colonists.

At the same time every zoologist will be willing to concede, that even if the departure of each species from a single centre had not appeared to be part of the plan of Nature, the range of species in general must have become limited, under the influence of a variety of causes, especially in the class of terrestrial mammalia. Scarcely any one of these could be expected to retain as fair a claim to the title of cosmopolite as man, although even the human race, fitted as it is by its bodily constitution and intellectual resources to spread very widely over the earth, is far from being strictly cosmopolite. It is excluded both from the arctic and antarctic circles, from many a wide desert and the summits of many mountain-chains; and lastly, from three-fourths of the globe covered by water, where there are large areas very prolific in animal life, even in the highest order of the vertebrate class. But the _habitations_ of species are, as before stated, in reference to plants (see above, p. 614), circumscribed by causes different from those which determine their _stations_, and these causes are clearly connected with the time and place of the original creation of each species.

As the names and characters of land quadrupeds are much better known to the general reader than those of other great families of the animal kingdom, I shall select this class to exemplify the zoological provinces into which species are divisible, confining myself, however, to those facts which may help to elucidate some principle, or rule apparently followed by the Author of Nature, in regard to that "mystery of mysteries," the first peopling of the earth with living beings.[873] First, then, the _European region_ comprehends, besides Europe, the borders of the Mediterranean, and even the north of Africa, and extends into Asia, beyond the Oural mountains and the Caspian. Although the species are almost all peculiar, the number of characteristic _genera_ is remarkably small. The bear, the fox, the hare, the rabbit, the deer, and almost every European form is found equally in several of the other large provinces of mammalia, where the species are distinct. Even the mole (_Talpa_), although confined to the northern parts of the old world, ranges eastwards, as far as the Himalaya mountains.

2dly. The _African_ Fauna, on the other hand, is singularly rich in generic forms, not met with in a living state in any other region. The hippopotamus, for example, of which two very distinct species are known, the giraffe, the Chimpanzee, the blue-faced baboon, the four-fingered monkeys (_Colubus_), many carnivora, such as _Proteles_, allied to the hyaena, and a multitude of other forms, are exclusively African. A few of the species inhabiting the northern confines of this continent, such as the dromedary, lion, and jackall, are also common to Asia; and a much larger number of _forms_ belong equally to the great Asiatic province, the species being distinct. The elephant, for example, of Africa is smaller, has a rounder head, and larger ears than the Indian one, and has only three instead of four nails on each hind foot. In like manner, not one of three African species of Rhinoceros agrees with one of the three Indian kinds.

3dly. The _Southern_ region of _Africa_, where that continent extends into the temperate zone, constitutes another separate zoological province, surrounded as it is on three sides by the ocean, and cut off from the countries of milder climate in the northern hemisphere, by the intervening torrid zone. In many instances, this region contains the same genera which are found in temperate climates to the northward of the line: but then the southern are different from the northern species. Thus, in the south we find the quagga and the zebra; in the north, the horse, the ass, and the jiggetai of Asia.

The south of Africa is spread out into fine level plains from the tropic to the Cape. In this region, says Pennant, besides the horse genus, of which five species have been found, there are also peculiar species of rhinoceros, the hog, and the hyrax, among pachydermatous races; and amongst the ruminating, the Cape buffalo, and a variety of remarkable antelopes, as the springbok, the oryx, the gnou, the leucophoa, the pygarga, and several others.[874]

4thly. The assemblage of quadrupeds in _Madagascar_ affords a striking illustration of the laws before alluded to, as governing the distribution of species in islands. Separated from Africa by the Mozambique channel, which is 300 miles wide, Madagascar forms, with two or three small islands in its immediate vicinity, a zoological province by itself, all the species except one, and nearly all the genera, being peculiar. The only exception consists of a small insectivorous quadruped (_Centetes_), found also in the Mauritius, to which place it is supposed to have been taken in ships. The most characteristic feature of this remarkable fauna consists in the number of quadrumana of the Lemur family, no less than six genera of these monkeys being exclusively met with in this island, and a seventh genus of the same, called _Galago_, which alone has any foreign representative, being found, as we might from analogy have anticipated, in the nearest main land. Had the species of quadrupeds in Madagascar agreed with those of the contiguous parts of Africa, as do those of England with the rest of Europe, the naturalist would have inferred that there had been a land communication since the period of the coming in of the existing quadrupeds, whereas we may now conclude that the Mozambique channel has constituted an insuperable barrier to the fusion of the continental fauna with that of the great island during the whole period that has elapsed since the living species were created.

5thly. Another of the great nations of terrestrial mammalia is that of _India_, containing a great variety of peculiar forms, such as the sloth-bear (_Prochilus_), the musk-deer (_Moscus_), the nylghau, the gibbon or long-armed ape, and many others.

6thly. A portion of the islands of the _Indian archipelago_ might, perhaps, be considered by some geologists as an appendage of the same province. In fact, we find in the large islands of Java, Sumatra, and Borneo, the same genera, for the most part, as on the continent of India, and some of the same species, _e. g._ the tapir (_Tapirus Malayanus_), the rhinoceros of Sumatra, and some others. Most of the species, however, are distinct, and each island has many, and even a few genera, peculiar to itself. Between eighty and ninety species are known to inhabit Java, and nearly the same number occur in Sumatra. Of these, more than half are common to the two islands. Borneo, which is much less explored, has yielded already upwards of sixty species, more than half of which are met with either in Java or Sumatra. Of the species inhabiting Sumatra and not found in Java, Borneo contains the greater portion. Upon the whole, if these three large islands were united, and a fusion of their respective indigenous mammalia should take place, they would present a fauna related to that of continental India, and comprising about as many species as we might expect from analogy to discover in an area of equal extent. The Philippine Islands are peopled with another assemblage of species generically related to the great Indian type.

7thly. But the islands of Celebes, Amboina, Timor, and _New Guinea_, constitute a different region of mammalia more allied to the Australian type, as having an intermixture of marsupial quadrupeds, yet showing an affinity also to the Indian in such forms as the deer (_Cervus_), the weasel (_Viverra_), the pig (_Sus_), the Macaque monkey (_Cercopithecus_), and others. As we proceed in a south-westerly direction, from Celebes to Amboina and thence to New Guinea, we find the Indian types diminishing in number, and the Australian (_i. e._ marsupial forms) increasing. Thus in New Guinea seven species of pouched quadrupeds have been detected, and among them two singular tree-kangaroos; yet only one species of the whole seven, viz. the flying opossum (_Petauris ariel_), is common to the Indian archipelago and the main land of Australia. The greater the zoological affinity, therefore, between the latter and the New Guinea fauna, although it seems in some way connected with geographical proximity, is not to be explained simply by the mutual migration of species from the one to the other.

8thly. When _Australia_ was discovered, its land quadrupeds, belonging almost exclusively to the marsupial or pouched tribe, such as the kangaroos, wombats, flying opossums, kangaroo-rats, and others, some feeding on herbs and fruits, others carnivorous, were so novel in their structure and aspect, that they appeared to the naturalist almost as strange as if they were the inhabitants of some other planet. We learn from the recent investigations of Mr. Waterhouse,[875] that no less than 170 species of marsupial quadrupeds have now been determined, and of the whole number all but thirty-two are exclusively restricted to Australia. Of these thirty-two, nine belong to the islands in the Indian archipelago before mentioned, and the other twenty-three are all species of opossum inhabiting the tropical parts of South America, or a few of them extending into Mexico and California, and one, the Virginian opossum, into the United States.

9thly. It only remains for me to say something of the mammiferous fauna of _North_ and _South America_. It has often been said that, where the three continents of Asia, Europe, and North America, approach very near to each other towards the pole, the whole arctic region forms one zoological and botanical province. The narrow straits which separate the old and new world are frozen over in winter, and the distance is farther lessened by intervening islands. Many plants and animals of various classes have accordingly spread over all the arctic lands, being sometimes carried in the same manner as the polar bear, when it is drifted on floating ice from Greenland to Iceland. But on a close inspection of the arctic mammalia, it has been found of late years that a very small number of the American species are identical with those of Europe or Asia. The genera are, in great part, the same or nearly allied; but the species are rarely identical, and are often very unlike, as in the case of the American badger and that of Europe. Some of the _genera_ of arctic America, such as the musk ox (_Ovibos_), are quite peculiar, and the distinctness of the fauna of the great continents goes on increasing in proportion as we trace them southwards, or as they recede farther from each other, and become more and more separated by the ocean. At length we find that the three groups of tropical mammalia, belonging severally to America, Africa, and India, have not a single species in common.

The predominant influence of climate over all the other causes which limit the range of species in the mammalia is perhaps nowhere so conspicuously displayed as in North America. The arctic fauna, so admirably described by Sir John Richardson, has scarcely any species in common with the fauna of the state of New York, which is 600 miles farther south, and comprises about forty distinct mammifers. If again we travel farther south about 600 miles, and enter another zone, running east and west, in South Carolina, Georgia, Alabama, and the contiguous states, we again meet with a new assemblage of land quadrupeds, and this again differs from the fauna of Texas, where frosts are unknown. It will be observed that on this continent there are no great geographical barriers running east and west, such as high snow-clad mountains, barren deserts, or wide arms of the sea, capable of checking the free migration of species from north to south. But notwithstanding the distinctness of those zones of indigenous mammalia, there are some species, such as the buffalo (_Bison Americanus_), the racoon (_Procyon lotor_), and the Virginian opossum (_Didelphis Virginiana_), which have a wider habitation, ranging almost from Canada to the Gulf of Mexico; but they form exceptions to the general rule. The opossum of Texas (_Didelphis carnivora_) is different from that of Virginia, and other species of the same genus inhabit westward of the Rocky Mountains, in California, for example, where almost all the mammalia differ from those of the United States.

10thly. The _West Indian_ land quadrupeds are not numerous, but several of them are peculiar; and 11thly, _South America_ is the most distinct, with the exception of Australia, of all the provinces into which the mammalia can be classed geographically. The various genera of monkeys, for example, belong to the family Platyrrhini, a large natural division of the quadrumana, so named from their widely separated nostrils. They have a peculiar dentition, and many of them prehensile tails, and are entirely unknown in other quarters of the globe. The sloths and armadillos, the true blood-sucking bats or vampyres (_Phyllostomidae_), the capybara, the largest of the rodents, the carnivorous coatimondi (_Nasua_), and a great many other forms, are also exclusively characteristic of South America.

"In Peru and Chili," says Humboldt, "the region of the grasses, which is at an elevation of from 12,300 to 15,400 feet, is inhabited by crowds of lama, guanaco, and alpaca. These quadrupeds, which here represent the genus camel of the ancient continent, have not extended themselves either to Brazil or Mexico; because, during their journey, they must necessarily have descended into regions that were too hot for them."[876] In this passage it will be seen that the doctrine of "specific centres" is tacitly assumed.

_Quadrupeds in Islands._--Islands remote from continents, especially those of small size, are either destitute of quadrupeds, except such as have been conveyed to them by man, or contain species peculiar to them. In the Galapagos archipelago no indigenous quadrupeds were found except one mouse, which is supposed to be distinct from any hitherto found elsewhere. A peculiar species of fox is indigenous in the Falkland Islands, and a rat in New Zealand, which last country, notwithstanding its magnitude, is destitute of other mammalia, except bats, and these, says Dr. Prichard, may have made their way along the chain of islands which extend from the shores of New Guinea far into the Southern Pacific. The same author remarks, that among the various groups of fertile islands in the Pacific, no quadrupeds have been met with except the rat and a few bats as above mentioned, and the dog and hog, which appear to have been conveyed thither by the natives from New Guinea. "Rats are to be found even on some desert islands, whither they may have been conveyed by canoes which have occasionally approached the shore. It is known, also, that rats occasionally swim in large numbers to considerable distances."[877]

_Geographical range of the Cetacea._--It is natural to suppose that the geographical range of the different species of Cetacea should be less correctly ascertained than that of the terrestrial mammifers. It is, however, well known that the whales which are obtained by our fishers in the South Seas are distinct from those of the North; and the same dissimilarity has been found in all the other marine animals, of the same class, so far as they have yet been studied by naturalists.

_Dispersion of quadrupeds._--Let us now inquire what facilities the various land quadrupeds enjoy of spreading themselves over the surface of the earth. In the first place, as their numbers multiply, all of them, whether they feed on plants, or prey on other animals, are disposed to scatter themselves gradually over as wide an area as is accessible to them. But before they have extended their migrations over a large space, they are usually arrested either by the sea, or a zone of uncongenial climate, or some lofty and unbroken chain of mountains, or a tract already occupied by a hostile and more powerful species.

_Their powers of swimming._--Rivers and narrow friths can seldom interfere with their progress; for the greater part of them swim well, and few are without this power when urged by danger and pressing want. Thus, amongst beasts of prey, the tiger is seen swimming about among the islands and creeks in the delta of the Ganges, and the jaguar traverses with ease the largest streams in South America.[878] The bear, also, and the bison, cross the current of the Mississippi. The popular error, that the common swine cannot escape by swimming when thrown into the water, has been contradicted by several curious and well-authenticated instances during the floods in Scotland of 1829. One pig, only six months old, after having been carried down from Garmouth to the bar at the mouth of the Spey, a distance of a quarter of a mile, swam four miles eastward to Port Gordon, and landed safe. Three others, of the same age and litter, swam, at the same time, five miles to the west, and landed at Blackhill.[879]

In an adult and wild state, these animals would doubtless have been more strong and active, and might, when hard pressed, have performed a much longer voyage. Hence islands remote from the continent may obtain inhabitants by casualties which, like the late storms in Morayshire, may only occur once in many centuries, or thousands of years, under all the same circumstances. It is obvious that powerful tides, winds, and currents may sometimes carry along quadrupeds capable, in like manner, of preserving themselves for hours in the sea, to very considerable distances; and in this way, perhaps, the tapir (_Tapir Indicus_) may have become common to Sumatra and the Malayan peninsula.

To the elephant, in particular, the power of crossing rivers is essential in a wild state, for the quantity of food which a herd of these animals consumes renders it necessary that they should be constantly moving from place to place. The elephant crosses the stream in two ways. If the bed of the river be hard, and the water not of too great a depth, he fords it. But when he crosses great rivers, such as the Ganges and the Niger, the elephant swims deep, so deep, that the end of his trunk only is out of the water; for it is a matter of indifference to him whether his body be completely immersed, provided he can bring the tip of his trunk to the surface, so as to breathe the external air.

Animals of the deer kind frequently take to the water, especially in the rutting season, when the stags are seen, swimming for several leagues at a time, from island to island, in search of the does, especially in the Canadian lakes; and in some countries where there are islands near the sea-shore, they fearlessly enter the sea and swim to them. In hunting excursions, in North America, the elk of that country is frequently pursued for great distances through the water.

The large herbivorous animals, which are gregarious, can never remain long in a confined region, as they consume so much vegetable food. The immense herds of bisons (_Bos Americanus_) which often, in the great valleys of the Mississippi and its tributaries blacken the surface of the prairie lands, are continually shifting their quarters, followed by wolves, which prowl about in their rear. "It is no exaggeration," says Mr. James, "to assert, that in one place, on the banks of the Platte, at least ten thousand bisons burst on our sight in an instant. In the morning, we again sought the living picture; but upon all the plain, which last evening was so teeming with noble animals, not one remained."[880]

_Migratory instincts._--Besides the disposition common to the individuals of every species slowly to extend their range in search of food, in proportion as their numbers augment, a migratory instinct often developes itself in an extraordinary manner, when, after an unusually prolific season, or upon a sudden scarcity of provisions, great multitudes are threatened by famine. It may be useful to enumerate some examples of these migrations, because they may put us upon our guard against attributing a high antiquity to a particular species merely because it is diffused over a great space; they show clearly how soon, in a state of nature, a newly created species might spread itself, in every direction, from a single point.

In very severe winters, great numbers of the black bears of America migrate from Canada into the United States; but in milder seasons, when they have been well fed, they remain and hybernate in the north.[881] The rein-deer, which, in Scandinavia, can scarcely exist to the south of the sixty-fifth parallel, descends, in consequence of the greater coldness of the climate, to the fiftieth degree in Chinese Tartary, and often roves into a country of more southern latitude than any part of England.

In Lapland, and other high latitudes, the common squirrels, whenever they are compelled, by want of provisions, to quit their usual abodes, migrate in amazing numbers, and travel directly forwards, allowing neither rocks nor forests, nor the broadest waters, to turn them from their course. Great numbers are often drowned in attempting to pass friths and rivers. In like manner the small Norway rat sometimes pursues its migrations in a straight line across rivers and lakes; and Pennant informs us, that when the rats, in Kamtschatka, become too numerous, they gather together in the spring, and proceed in great bodies westward, swimming over rivers, lakes, and arms of the sea. Many are drowned or destroyed by water-fowl or fish. As soon as they have crossed the river Penginsk, at the head of the gulf of the same name, they turn southward, and reach the rivers Judoma and Okotsk by the middle of July; a district more than 800 miles distant from their point of departure.

The lemings, also, a small kind of rat, are described as natives of the mountains of Kolen, in Lapland; and once or twice in a quarter of a century they appear in vast numbers, advancing along the ground, and "devouring every green thing." Innumerable bands march from the Kolen, through Nordland and Finmark, to the Western Ocean, which they immediately enter; and after swimming about for some time, perish. Other bands take their route through Swedish Lapland, to the Bothnian Gulf, where they are drowned in the same manner. They are followed in their journeys by bears, wolves, and foxes, which prey upon them incessantly. They generally move in lines, which are about three feet from each other, and exactly parallel, going directly forward through rivers and lakes; and when they meet with stacks of hay or corn, gnawing their way through them instead of passing round.[882] These excursions usually precede a rigorous winter, of which the lemings seem in some way forewarned.

The Leming, or Lapland Marmot (Mus Lemmus, Linn.)]

Vast troops of the wild ass, or _onager_ of the ancients, which inhabit the mountainous deserts of Great Tartary, feed, during the summer, in the tracts east and north of Lake Aral. In the autumn they collect in herds of hundreds, and even thousands, and direct their course towards Persia, to enjoy a warm retreat during winter.[883] Bands of two or three hundred quaggas, a species of wild ass, are sometimes seen to migrate from the tropical plains of southern Africa to the vicinity of the Malaleveen River. During their migrations they are followed by lions, who slaughter them night by night.[884]

The migratory swarms of the springbok, or Cape antelope, afford another illustration of the rapidity with which a species under certain circumstances may be diffused over a continent. When the stagnant pools of the immense deserts south of the Orange River dry up, which often happens after intervals of three or four years, myriads of these animals desert the parched soil, and pour down like a deluge on the cultivated regions near the Cape. The havoc committed by them resembles that of the African locusts; and so crowded are the herds, that "the lion has been seen to walk in the midst of the compressed phalanx with only as much room between him and his victims as the fears of those immediately around could procure by pressing outwards."[885]

Mydaus meliceps, or badger-headed Mydaus. Length, including the tail, 16 inches.]

Dr. Horsfield mentions a singular fact in regard to the geographical distribution of the _Mydaus meliceps_, an animal intermediate between the polecat and badger. It inhabits Java, and is "confined exclusively to those mountains which have an elevation of more than seven thousand feet above the level of the ocean; on these it occurs with the same regularity as many plants. The long extended surface of Java, abounding with conical points which exceed this elevation, affords many places favorable for its resort. On ascending these mountains, the traveller scarcely fails to meet with this animal, which, from its peculiarities, is universally known to the inhabitants of these elevated tracts, while to those of the plains it is as strange to an animal from a foreign county. In my visits to the mountainous districts, I uniformly met with it; and, as far as the information of the natives can be relied on, it is found on all the mountains."[886]

Now, if asked to conjecture how the Mydaus arrived at the elevated regions of each of these isolated mountains, we might say that, before the island was peopled by man, by whom their numbers are now thinned, they may occasionally have multiplied so as to be forced to collect together and migrate: in which case notwithstanding the slowness of their motions, some few would succeed in reaching another mountain, some twenty, or even, perhaps, fifty miles distant; for although the climate of the hot intervening plains would be unfavourable to them, they might support it for a time, and would find there abundance of insects on which they feed. Volcanic eruptions, which, at different times have covered the summits of some of those lofty cones with sterile sand and ashes, may have occasionally contributed to force on these migrations.

_Drifting of animals on ice-floes._--The power of the terrestrial mammalia to cross the sea is very limited, and it was before stated that the same species is scarcely ever common to districts widely separated by the ocean. If there be some exceptions to this rule, they generally admit of explanation; for there are natural means whereby some animals may be floated across the water, and the sea may in the course of ages wear a wide passage through a neck of land, leaving individuals of a species on each side of the new channel. Polar bears are known to have been frequently drifted on the ice from Greenland to Iceland; they can also swim to considerable distances, for Captain Parry, on the return of his ships through Barrow's Straits, met with a bear swimming in the water about midway between the shores, which were about forty miles apart, and where no ice was in sight.[887] "Near the east coast of Greenland," observes Scoresby, "they have been seen on the ice in such quantities, that they were compared to flocks of sheep on a common; and they are often found on field-ice, above two hundred miles from the shore."[888] Wolves, in the arctic regions, often venture upon the ice near the shore, for the purpose of preying upon young seals which they surprise when asleep. When these ice-floes get detached, the wolves are often carried out to sea; and though some may be drifted to islands or continents, the greater part of them perish, and have been often heard in this situation howling dreadfully, as they die by famine.[889]

During the short summer which visits Melville Island, various plants push forth their leaves and flowers the moment the snow is off the ground, and form a carpet spangled with the most lively colours. These secluded spots are reached annually by herds of musk-oxen and reindeer, which travel immense distances over dreary and desolate regions, to graze undisturbed on these luxuriant pastures.[890] The rein-deer often pass along in the same manner, by the chain of the Aleutian Islands, from Behring's Straits to Kamtschatka, subsisting on the moss found in these islands during their passage.[891] But the musk-ox, notwithstanding its migratory habits, and its long journeys over the ice, does not exist, either in Asia or Greenland.[892]

_On floating islands of drift-wood._--Within the tropics there are no ice-floes; but, as if to compensate for that mode of transportation, there are floating islets of matted trees, which are often borne along through considerable spaces. These are sometimes seen sailing at the distance of fifty or one hundred miles from the mouth of the Ganges, with living trees standing erect upon them. The Amazon, the Congo, and the Orinoco, also produce these verdant rafts, which are formed in the manner already described when speaking of the great raft of the Atchafalaya, an arm of the Mississippi, where a natural bridge of timber, ten miles long, and more than two hundred yards wide, existed for more than forty years, supporting a luxuriant vegetation, and rising and sinking with the water which flowed beneath it.

On these green islets of the Mississippi, observes Malte-Brun, young trees take root, and the pistia and nenuphar display their yellow flowers: serpents, birds, and the cayman alligator, come to repose there, and all are sometimes carried to the sea and engulphed in its waters.[893]

Spix and Martius relate that, during their travels in Brazil, they were exposed to great danger while ascending the Amazon in a canoe, from the vast quantity of drift-wood constantly propelled against them by the current; so much so, that their safety depended on the crew being always on the alert to turn aside the trunks of trees with long poles. The tops alone of some trees appeared above water, others had their roots attached to them with so much soil that they might be compared to floating islets. On these, say the travellers, we saw some very singular assemblages of animals, pursuing peacefully their uncertain way in strange companionship. On one raft were several grave-looking storks, perched by the side of a party of monkeys, who made comical gestures, and burst into loud cries, on seeing the canoe. On another was seen a number of ducks and divers, sitting by a group of squirrels. Next came down upon the stem of a large rotten cedar tree, an enormous crocodile, by the side of a tiger-cat, both animals regarding each other with hostility and mistrust, but the saurian being evidently most at his ease, as conscious of his superior strength.[894]

Similar green rafts, principally composed of canes and brushwood, are called "camelotes" on the Parana in South America; and they are occasionally carried down by inundations, bearing on them the tiger, cayman, squirrels, and other quadrupeds, which are said to be always terror-stricken on their floating habitation. No less than four tigers (pumas) were landed in this manner in one night at Monte Video, lat. 35 degrees S., to the great alarm of the inhabitants, who found them prowling about the streets in the morning.[895]

In a memoir lately published, a naval officer relates that, as he returned from China by the eastern passage, he fell in, among the Moluccas, with several small floating islands of this kind, covered with mangrove trees interwoven with underwood. The trees and shrubs retained their verdure, receiving nourishment from a stratum of soil which formed a white beach round the margin of each raft, where it was exposed to the washing of the waves and the rays of the sun.[896] The occurrence of soil in such situations may easily be explained; for all the natural bridges of timber which occasionally connect the islands of the Ganges, Mississippi, and other rivers, with their banks, are exposed to floods of water, densely charged with sediment.

Captain W. H. Smyth informs me, that, when cruising in the Cornwallis amidst the Philippine Islands, he has more than once seen, after those dreadful hurricanes called typhoons, floating masses of wood, with trees growing upon them, and ships have sometimes been in imminent peril, as often as these islands were mistaken for terra firma, when, in fact, they were in rapid motion.

It is highly interesting to trace, in imagination, the effects of the passage of these rafts from the mouth of a large river to some archipelago, such as those in the South Pacific, raised from the deep, in comparatively modern times, by the operations of the volcano and the earthquake, and the joint labours of coral animals and testacea. If a storm arise, and the frail vessel be wrecked, still many a bird and insect may succeed in gaining, by flight, some island of the newly formed group, while the seeds and berries of herbs and shrubs, which fall into the waves, may be thrown upon the strand. But if the surface of the deep be calm, and the rafts are carried along by a current, or wafted by some slight breath of air fanning the foliage of the green trees, it may arrive, after a passage of several weeks, at the bay of an island, into which its plants and animals may be poured out as from an ark, and thus a colony of several hundred new species may at once be naturalized.

The reader should be reminded, that I merely advert to the transportation of these rafts as of extremely rare and accidental occurrence; but it may account, in tropical countries, for some of the rare exceptions to the general law of the confined range of mammiferous species.

_Migrations of the Cetacea._--Many of the Cetacea, the whales of the northern seas for example, are found to desert one tract of the sea, and to visit another very distant, when they are urged by want of food, or danger. The seals also retire from the coast of Greenland in July, return again in September, and depart again in March, to return in June. They proceed in great droves northwards, directing their course where the sea is most free from ice, and are observed to be extremely fat when they set out on this expedition, and very lean when they come home again.[897]

_Species of the Mediterranean, Black Sea, and Caspian identical._--Some naturalists have wondered that the sea-calves, dolphins, and other marine mammalia of the Mediterranean and Black Sea, should be identical with those found in the Caspian: and among other fanciful theories, they have suggested that they may dive through subterranean conduits, and thus pass from one sea into the other. But as the occurrence of wolves and other noxious animals, on both sides of the British Channel, was adduced, by Verstegan and Desmarest, as one of many arguments to prove that England and France were once united; so the correspondence of the aquatic species of the inland seas of Asia with those of the Black Sea tend to confirm the hypothesis, for which there are abundance of independent geological data, that those seas were connected together by straits at no remote period of the earth's history.

_Geographical Distribution and Migrations of Birds._

I shall now offer a few observations on some of the other divisions of the animal kingdom. Birds, notwithstanding their great locomotive powers, form no exception to the general rules already laid down; but, in this class, as in plants and terrestrial quadrupeds, different groups of species are circumscribed within definite limits. We find, for example, one assemblage in the Brazils, another in the same latitudes in Central Africa, another in India, and a fourth in New Holland. Of twenty-six different species of land birds found in the Galapagos archipelago, all, with the exception of one, are distinct from those inhabiting other parts of the globe;[898] and in other archipelagos a single island sometimes contains a species found in no other spot on the whole earth; as is exemplified in some of the parrot tribes. In this extensive family, which are, with few exceptions, inhabitants of tropical regions, the American group has not one in common with the African, nor either of these with the parrots of India.[899]

Another illustration is afforded by that minute and beautiful tribe, the humming-birds. The whole of them are, in the first place, peculiar to the new world; but some species are confined to Mexico, while others exist only in some of the West India Islands, and have not been found elsewhere in the western hemisphere. Yet there are species of this family which have a vast range, as the _Trochilus flammifrons_ (or _Mellisuga Kingii_), which is found over a space of 2500 miles on the west coast of South America, from the hot dry country of Lima to the humid forests of Tierra del Fuego. Captain King, during his survey in the years 1826-30, found this bird at the Straits of Magellan, in the month of May--the depth of winter--sucking the flowers of a large species of fuchsia, then in bloom, in the midst of a shower of snow.

The ornithology of our own country affords one well-known and striking exemplification of the law of a limited specific range; for the common grouse (_Tetra scoticus_) occurs nowhere in the known world except in the British isles.

Some species of the vulture tribe are said to be cosmopolites; and the common wild goose (_Anas anser_, Linn.), if we may believe some ornithologists, is a general inhabitant of the globe, being met with from Lapland to the Cape of Good Hope, frequent in Arabia, Persia, China, and Japan, and in the American continent from Hudson's Bay to South Carolina.[900] An extraordinary range has also been attributed to the nightingale, which extends from western Europe to Persia, and still farther. In a work entitled Specchio Comparativo,[901] by Charles Bonaparte, many species of birds are enumerated as common to Rome and Philadelphia: the greater part of these are migratory, but some of them, such as the long-eared owl (_Strix otus_), are permanent in both countries. The correspondence of the ornithological fauna of the eastern and western hemispheres increases considerably, as might have been anticipated, in high northern latitudes.[902]

_Their facilities of diffusion._--In parallel zones of the northern and southern hemispheres, a great general correspondence of form is observable, both in the aquatic and terrestrial birds; but there is rarely any specific identity; and this phenomenon is truly remarkable, when we recollect the readiness with which some birds, not gifted with great powers of flight, shift their quarters to different regions, and the facility with which others, possessing great strength of wing, perform their aarial voyage. Some migrate periodically from high latitudes, to avoid the cold of winter, and the accompaniments of cold,--scarcity of insects and vegetable food; others, it is said, for some particular kinds of nutriment required for rearing their young: for this purpose they often traverse the ocean for thousands of miles, and recross it at other periods, with equal security.

Periodical migrations, no less regular, are mentioned by Humboldt, of many American water-fowl, from one part of the tropics to another, in a zone where there is the same temperature throughout the year. Immense flights of ducks leave the valley of the Orinoco, when the increasing depth of its waters and the flooding of its shores prevent them from catching fish, insects, and aquatic worms. They then betake themselves to the Rio Negro and Amazon, having passed from the eighth and third degrees of north latitude to the first and fourth of south latitude, directing their course south-south-east. In September, when the Orinoco decreases and re-enters its channel, these birds return northwards.[903]

The insectivorous swallows which visit our island would perish during winter, if they did not annually repair to warmer climes. It is supposed that in these aerial excursions the average rapidity of their flight is not less than fifty miles an hour; so that, when aided by the wind, they soon reach warmer latitudes. Spallanzani calculated that the swallow can fly at the rate of ninety-two miles an hour, and conceived that the rapidity of the swift might be three times greater.[904] The rate of flight of the eider duck (_Anas mollissima_) is said to be ninety miles an hour; and Bachman says that the hawk, wild pigeon (_Columba migratoria_), and several species of wild ducks, in North America, fly at the rate of forty miles an hour, or nearly a thousand miles in twenty-four hours.[905]

When we reflect how easily different species, in a great lapse of ages, may be each overtaken by gales and hurricanes, and, abandoning themselves to the tempest, be scattered at random through various regions of the earth's surface, where the temperature of the atmosphere, the vegetation, and the animal productions, might be suited to their wants, we shall be prepared to find some species capriciously distributed, and to be sometimes unable to determine the native countries of each. Captain Smyth informs me, that, when engaged in his survey of the Mediterranean, he encountered a gale in the Gulf of Lyons, at the distance of between twenty and thirty leagues from the coast of France, which bore along many land birds of various species, some of which alighted on the ship, while others were thrown with violence against the sails. In this manner islands become tenanted by species of birds inhabiting the nearest mainland.

_Geographical Distribution and Dissemination of Reptiles._

A few facts respecting the third great class of vertebrated animals will suffice to show that the plan of nature in regard to their location on the globe is perfectly analogous to that already exemplified in other parts of the organic creation, and has probably been determined by similar causes.

_Habitations of reptiles._--Of the great saurians, the gavials which inhabit the Ganges differ from the cayman of America, or the crocodile of the Nile. The monitor of New Holland is specifically distinct from the Indian species; these latter, again, from the African, and all from their congeners in the new world. So in regard to snakes; we find the boa of America represented by the python, a different though nearly allied genus in India. America is the country of the rattlesnake; Africa, of the cerastes; and Asia, of the hooded snake, or cobra di capello. The amphibious genera Siren and Menopoma belong to North America, possessing both lungs and gills, and respiring at pleasure either air or water. The only analogous animal of the old world is the _Proteus anguinus_ of the lakes of Lower Carniola, and the grotto of Adelsberg between Trieste and Vienna.[906]

There is a legend that St. Patrick expelled all reptiles from Ireland; and certain it is that none of the three species of snakes common in England, nor the toad, have been observed there by naturalists. They have our common frog, and our water-newt, and according to Ray (Quad. 264.), the green lizard (_Lacerta viridis_).

_Migrations of the larger reptiles._--The range of the large reptiles is, in general, quite as limited as that of some orders of the terrestrial mammalia. The great saurians sometimes cross a considerable tract in order to pass from one river to another; but their motions by land are generally slower than those of quadrupeds. By water, however, they may transport themselves to distant situations more easily. The larger alligator of the Ganges sometimes descends beyond the brackish water of the delta into the sea; and in such cases it might chance to be drifted away by a current, and survive till it reached a shore at some distance; but such casualties are probably very rare.

Turtles migrate in large droves from one part of the ocean to another during the ovipositing season; and they find their way annually to the island of Ascension, from which the nearest land is about 800 miles distant. Dr. Fleming mentions, that an individual of the hawk's bill turtle (_Chelonia imbricata_), so common in the American seas, has been taken at Papa Stour, one of the West Zetland Islands;[907] and, according to Sibbald, "the same animal came into Orkney." Another was taken, in 1774, in the Severn, according to Turton. Two instances, also, of the occurrence of the leathern tortoise (_C. coriacea_), on the coast of Cornwall, in 1756, are mentioned by Borlase. These animals of more southern seas can be considered only as stragglers, attracted to our shores during uncommonly warm seasons by an abundant supply of food, or carried by the Gulf stream, or driven by storms to high latitudes.

Some of the smaller reptiles lay their eggs on aquatic plants; and these must often be borne rapidly by rivers, and conveyed to distant regions in a manner similar to the dispersion of seeds before adverted to. But that the larger ophidians may be themselves transported across the seas, is evident from the following most interesting account of the arrival of one at the island of St. Vincent. It is worthy of being recorded, says Mr. Guilding, "that a noble specimen of the _Boa constrictor_ was lately conveyed to us by the currents, twisted round the trunk of a large sound cedar tree, which had probably been washed out of the bank by the floods of some great South American river, while its huge folds hung on the branches, as it waited for its prey. The monster was fortunately destroyed after killing a few sheep, and his skeleton now hangs before me in my study, putting me in mind how much reason I might have had to fear in my future rambles through the forests of St. Vincent, had this formidable reptile been a pregnant female, and escaped to a safe retreat."[908]