Chapter IV: Part 4

=Distribution.=--Nova Scotia to Manitoba, south to Georgia and west to Texas and Wyoming. Found in all parts of Indiana, although we have no reports from the knob area where no doubt it is only local. It is a tree of wet woods, low borders of streams, etc., except among the hills of southern Indiana, it is an occasional tree of the slopes. In favorable habitats it was a frequent to a common tree. Its most constant associates are white elm, swamp white and red oak, linn, green and black ash, shellbark hickory, etc. It is sometimes called mossy-cup oak.

=Remarks.=--Wood and uses similar to that of white oak. In point of number, size and value it ranks as one of the most valuable trees of the State. Michaux[39] says: "A tree three miles from Troy, Ohio, was measured that was fourteen feet and nine inches in diameter six feet above the ground. The trunk rises about fifty feet without limbs, and with scarcely a perceptible diminution in size."

=7a.= =Quercus macrocarpa= var. =olivæfórmis= (Michaux filius) Gray. This variety is distinguished from the typical form by its shallow cup, and the long oval nut which is often 3 cm. long. The cup is semi-hemispheric, and encloses the nut for about one-half its length.

Authentic specimens are at hand from Wells County, and it has been reported from Gibson and Hamilton Counties. No doubt this form has a wider range.

=8.= =Quercus lyràta= Walter. Overcup Oak. Plate 47. Medium sized trees; bark generally intermediate between that of the swamp white and bur oak; leaves on petioles 5-30 mm. long which are generally somewhat reddish toward the base, 10-20 cm. long, obovate or oblong-obovate, margins very irregularly divided into 5-9 short or long lobes, ascending and generally acute, ordinarily the three terminal lobes are the largest, base of leaves wedge-shape, or narrowly rounded, upper surface at maturity dark green and smooth, the under surface densely covered with a thick tomentum to which is added more or less long and single or fascicled straight hairs; when the leaves are as described on the under surface they are gray beneath; however, a form occurs which is yellow green beneath and has little or no tomentum, but is thickly covered with long single or fascicled straight hairs; acorn single or in pairs, on stalks generally about 1 cm. long, sometimes the stalks are 3 cm. long, the stalk lies in a plane at a right angle to the base of the acorn which is a characteristic of this species; nut depressed-globose, about 1.5 cm. long, generally almost completely enclosed in the cup, or sometimes enclosed only for about 2/3 its length; cup generally very thick at the base, gradually becoming thinner at the top, and often it splits open; scales tomentose on the back, those near the base, thick and tuberculate on the back and blunt, but those near the top of the cup are acute or long attenuate; kernel sweet.

=Distribution.=--Maryland to Missouri,[40] and south to Florida and west to Texas. In Indiana it is found only about river sloughs or deep swamps in the southwestern counties. At present it is known only from Knox, Gibson, Posey and Spencer Counties. It was reported by Nieuwland[41] for Marshall County on the authority of Clark. This specimen was taken during a survey of Lake Maxinkuckee, and is deposited in the National Museum. I have had the specimen examined by an authority, who reports that it is some other species. Its habitat is that of areas that are inundated much of the winter season. It is so rare that its associates could not be learned. In one place it grew in a depression lower than a nearby pin oak, and in another place it grew in a depression in a very low woods, surrounded by sweet gum, big shell bark hickory, and pin oak. It is generally found singly in depressions, but it is a common tree on the low border of the west side of Burnett's pond in Gibson County.

=Remarks.=--Wood and uses similar to that of white oak. In our area it is usually known as bur oak.

QUERCUS LYRATA Walter. Overcup Oak. (× 1/2.)

Acorns from different trees.]

=9.= =Quercus imbricària= Michaux. Shingle Oak. Plate 48. Medium to large sized trees; leaves on petioles generally 0.5-1 cm. long, 7-16 cm. long, elliptic to oblong-lanceolate, narrowed or rounded at the base, apex generally sharp-pointed and ending with a bristle, sometimes very wide leaves are blunt at the apex, margins entire, when they first appear the upper surface is somewhat woolly and the under surface whitish with a dense tomentum, soon glabrous and a dark green above, remaining more or less densely woolly or pubescent beneath; acorns sessile or nearly so, solitary or in pairs; nuts ovoid, about 1 cm. long and enclosed for about 1/2 their length in the cup; cup rounded at the base; scales pubescent on the back and obtuse.

=Distribution.=--Pennsylvania, Michigan to Nebraska, south to Georgia and west to Arkansas. Found throughout Indiana. It is essentially a tree of low ground, but it is sometimes found near the base of slopes, and in the knob area it is sometimes found on the crest of ridges. In all parts of Indiana except the southwestern part it is found only locally and then usually in colonies of a few trees. In Wells County, I know of only two trees located at the base of a slope bordering a pond in Jackson Township. In the southwestern part of the State it is frequent to a common tree in its peculiar habitat. It appears that when drainage basins decrease in size, and leave sandy river bottoms, and bordering low sand dunes, that the shingle oak is the first oak to occupy the area. On the sand ridges it is crowded out by the black, black jack and post oaks. In the bottoms it is succeeded by pin, Schneck's, Spanish, swamp white and post oaks. Special notes were made on its distribution on a trip through Gibson, Pike, Daviess, Greene and Sullivan Counties, going from Francisco northward through the Patoka bottoms where in many places it forms pure stands. Usually in situations a little higher than the pin oak zone. Thence eastward to Winslow and then north to Sandy Hook in Daviess County, thence north to Washington, Montgomery, Odon, Newberry, Lyons, Marco and Sullivan. In its habitat all along this route it was a frequent to a very common tree. A few miles northeast of Montgomery is a small area which a pioneer informed me was originally a prairie. Typical prairie plants are yet found along the roadside and fences in the area. I was informed that the shingle oak was the only species found on the area, and on the border of the area. It is believed the mass distribution of the species was in the area indicated by the preceding route. Both east and west of this area the species becomes less frequent.

=Remarks.=--Wood similar to red oak, but much inferior. Evidently it is rather a slow growing tree, but it might find a use as a shade or ornamental tree in sandy habitats where the pin oak would not thrive. It is also called black oak, peach oak, jack oak and water oak.

QUERCUS IMBRICARIA Michaux. Shingle Oak. (× 1/2.)]

=10.= =Quercus rùbra= Linnæus. [_Quercus maxima_ (Marshall) Ashe of some recent authors]. Red Oak. Plate 49. Large trees; winter buds ovoid, pointed, reddish, outer scales glabrous, sometimes pubescent on the edges; twigs soon smooth and reddish; leaves on petioles 2.5-5 cm. long, 10-20 cm. long, oval to oblong-obovate, broadly wedge-shape or truncate at the base, the margins divided by wide or narrow sinuses generally into 7-9 lobes, sometimes as many as 11, the lobes not uniform in size or shape, lobes broadest at the base and ending generally in 1-5 bristle points, pubescent above and below at first, soon becoming smooth at maturity and a dark green above, paler and yellowish-green beneath and smooth or with tufts of tomentum in the axils of the veins; acorns solitary or in pairs, sessile or on very short stalks; nuts ovoid, flat at the base, and rounded at the apex, 2-3 cm. long, enclosed for about 1/4 their length in the shallow cup; cups 2-3 cm. in diameter, thick, saucer-shape, flat or only slightly rounded at the base; scales ovate, blunt, appressed, and pubescent on the back; kernel somewhat bitter, eaten by hogs and cattle, but not relished by wild animals.

=Distribution.=--Nova Scotia to Minnesota, south to Florida and west to Texas. Found throughout Indiana, although local in the knob area. Its preferred habitat is that of moist, rich and fairly well drained woods. It does not thrive in situations that are inundated much of the winter season such as the pin oak will endure. In the southern part of the State, especially in the flats it is frequently found on the high bluffs of streams and very large forest trees are frequent on a dry wooded slope of ten acres, on the Davis farm four miles south of Salem. In a congenial habitat it was a frequent to a common tree, although such a thing as nearly a pure stand would never be met with, such as was often formed by the white, black, shingle or pin oak.

=Remarks.=--Wood hard, heavy, strong, close-grained, but not as good as white oak in any of its mechanical qualities. Commercially all of the biennial oaks are usually considered as red oak. The true red oak, however, is generally considered the best of all the biennial oaks. Until recently, when white oak became scarce, red oak was not in much demand, and was used principally for construction material. Now it is substituted in many places for white oak, and the uses now are in a great measure the same as those of white oak.

The red oak grows rapidly, and is able to adapt itself to many soil conditions. It has been used in European countries for two centuries for shade and ornamental planting. It reproduces easily by planting the acorns, and should receive attention by woodlot owners as a suitable species for reinforcing woodlands, or in general forest planting.

QUERCUS RUBRA Linnæus. Red Oak. (× 1/2.)

Acorns from different trees.]

=11.= =Quercus palústris= Du Roi. Pin Oak. Plate 50. Medium to large trees with very tight bark, the furrows shallow and generally wide; twigs at first pubescent, soon becoming smooth and reddish-brown; leaves on petioles generally 1-5 cm. long, blades about 7-15 cm. long, usually about 2/3 as wide, sometimes as wide as long, ovate to obovate in outline, narrowed to broadly truncate at the base, the margins divided into 5-7 lobes by deep and wide sinuses, except leaves that grow in the shade, the sinus cuts the blade to more than half way to the midrib, the lobes are widest at the base, or sometimes widest near the apex, the lobes usually somewhat toothed or lobed and end in 1-7 bristle tips, leaves hairy when they first appear, soon becoming glabrate and a glossy dark green above, a paler green beneath and smooth except tufts of hairs in the axils of the principal veins; acorns sessile or nearly so, single or in clusters; nuts subglobose or ovoid, generally 10-12 mm. long, the ovoid form somewhat smaller, covered about 1/4 their length by the shallow cups; cups saucer-shape and generally flat on the bottom, those with the ovoid nuts are rounded on the bottom; scales pubescent on the back, and rounded or blunt at the apex.

=Distribution.=--Massachusetts, southwestern Ontario, Michigan to Iowa and south to Virginia and west to Oklahoma. Found in every county of Indiana. It is found only in wet situations where it is a frequent to a common tree. It prefers a hard compact clay soil with little drainage hence is rarely met with on the low borders of lakes where the soil is principally organic matter.

=Remarks.=--Wood similar to red oak, but much inferior to it. It is tardy in the natural pruning of its lower branches, and when the dead branches break off they usually do so at some distance from the trunk. The stumps of the dead branches which penetrate to the center of the tree have given it the name of pin oak. It is also sometimes called water oak, and swamp oak.

For street and ornamental planting it is one of the most desirable oaks to use. It is adapted to a moist soil, grows rapidly, and produces a dense shade. When grown in the open it develops a pyramidal crown.

The nut of this species always has a depressed form, except a tree or two in Wells County which produce ovate nuts which are cone-pointed, and in bulk about half the size of the ordinary form. This form should be looked for to ascertain its area of distribution.

QUERCUS PALUSTRIS Muenchhausen. (× 1/2.)

Acorns from different trees. Those on the left the common form, those on the right the rare form.]

QUERCUS SCHNECKII Britton. Schneck's Oak. (× 1/2.)

Specimens from type tree.]

=12.= =Quercus Schnéckii= Britton. Schneck's Oak. Plate 51. Large trees; bark somewhat intermediate between pin and red oak; twigs gray by autumn; winter buds large, about 0.5 cm. long, ovoid, glabrous and gray; leaves on petioles 2-6 cm. long, blades generally 8-18 cm. long, generally truncate at the base, sometimes wedge-shaped, leaves ovate to obovate in outline, divided into 5-7 lobes, by deep rounded and wide sinuses, the sinuses cutting the blade to more than half way to the midrib, except the leaves of lower branches that grow in the shade, the lobes variable in shape and size, usually the lowest are the shortest and smaller, the middle the longest and largest, the lobes are sometimes widest at the base, and sometimes widest at the apex, the end of the lobes are more or less toothed or lobed; the leaves at maturity are bright green, glossy and smooth above, a paler and yellow green and smooth beneath except tufts of hairs in the axils of the principal veins; acorns solitary or in pairs, usually on stalks about 0.5 cm. long; nuts ovoid, sometimes broadly so, or oblong, broad and flat or slightly convex at base, usually 1.5-2 cm. long, enclosed in the cup from 1/4-1/3 their length; cups flat or convex at the base: scales generally pubescent on the back, gray or with a reddish tip on those of the Lower Wabash Valley, or reddish gray and with margins more or less red of trees of the Upper Wabash Valley.

=Distribution.=--In Indiana this species has been reported only from Wells, Bartholomew, Vermillion, Knox, Gibson and Posey Counties. This species was not separated from our common red oak until after all of the local floras of Indiana had been written, and it may have a much wider range than is at present known. In Wells County it is the prevailing "red oak" of the county, and no doubt is distributed throughout the Wabash Valley. In this area it is associated with all moist ground species. In the lower Wabash Valley, especially in Gibson, Knox and Posey Counties it is associated with Spanish, pin, and shingle oaks, sweet gum, etc. Several trees were noted in Knox County in Little Cypress swamp where it was associated with cypress, pin oak, white elm, red maple and swell-butt ash.

=Remarks.=--This anomalous red oak has a range from Indiana to Texas. When the attention of authors was directed to it, several new species were the result. Later authors are not agreed as to whether this form, which has such a wide range and hence liable to show considerable variation within such a long range, is one or several species. C. S. Sargent who for years has studied this form throughout its range has seen the author's specimens and calls those with shallow cups typical or nearly typical _Quercus Shumardii_ Buckley[42] and those with the deep cups _Quercus Shumardii_ variety _Schneckii_ (Britton) Sargent.

The writer has made rather an intensive study of the forms in Wells County and in the Lower Wabash Valley and has not been able to satisfy himself that, allowing for a reasonable variation, there is even a varietal difference in Indiana forms. The description has been drawn to cover all of the forms of Indiana.

Dr. J. Schneck of Mt. Carmel, Illinois, was one of the first to discover that this form was not our common red oak, and when he called Dr. Britton's attention to it, Dr. Britton named it _Quercus Schneckii_ in honor of its discoverer.

=13.= =Quercus ellipsoidàlis.= E. J. Hill. Hill's Oak. Plate 52. Medium sized trees; inner bark yellowish; twigs pubescent at first, becoming smooth and reddish brown by autumn; leaves on petioles 2-5 cm. long, ovate to slightly obovate or nearly orbicular in outline, 7-15 cm. long, wedge-shape or, truncate at the base, margin divided into 5-7 long lobes by wide sinuses which usually extend to more than half way to the midrib, sinuses rounded at the base, lobes broadest at the base or the apex, ending in 1-7 bristle points, leaves at first pubescent, both above and below, soon becoming glabrous above, and smooth beneath except tufts of hairs in the axils of the principal veins; acorns nearly sessile or on short stalks, single or in pairs; nuts oval to oblong, 12-20 mm. long, enclosed for 1/3-1/2 their length in the cup; scales obtuse, light reddish-brown, pubescent on the back; kernel pale yellow and bitter.

=Distribution.=--Northwestern Indiana to Manitoba and south to Iowa. In Indiana it has been reported only from Lake and Porter Counties by Hill, and from White County by Heimlich. According to Hill, who has made the most extensive study of the distribution of this species in our area, the tree is found on sandy and clayey uplands, and in moist sandy places. It closely resembles the pin oak for which it has been mistaken. It also resembles the black and scarlet oaks. We have very little data on the range or distribution of the species in this State.

=14.= =Quercus velùtina= Lamarck. Black Oak. Plate 53. Medium to large sized trees; inner bark yellow or orange; leaves on petioles 2-8 cm. long, ovate oblong or obovate, very variable in outline and in size, those of young trees and coppice shoots being very large, those of mature trees usually 12-18 cm. long, wedge-shape or truncate at the base, the margin divided into 5-9 lobes by wide and usually deep sinuses which are rounded at the base, the lobes variable in shape and size, the terminals of many of the lobes toothed or slightly lobed and ending in one or more bristles, leaves pubescent on both sides at first, soon becoming smooth, glossy and a dark green above; leaves of fruiting branches usually smooth beneath except the tufts of brown hairs in the axils of the principal veins, or rarely more or less pubescent over the whole under surface, the under surface of leaves of sterile branches and young trees usually are the most pubescent beneath, the leaves of some trees are much like those of the scarlet oak, but on the whole are larger; acorns sessile or nearly so, single or in pairs; nuts ovoid, oblong or subglobose, 1.5-2 cm. long, enclosed for about half their length in the cup-shaped cup; scales light-brown, densely pubescent on the back, obtuse, loose above the middle of the cup; kernel bitter.

QUERCUS ELLIPSOIDALIS E. J. Hill. Hill's Oak. (× 1/2.)

Specimens from type tree.]

QUERCUS VELUTINA Lamarck. Black Oak. (× 1/2.)]

=Distribution.=--Maine, southern Ontario, southern Minnesota, southern Nebraska south to Florida and west to Texas. Found throughout Indiana. It was no doubt found in every county or nearly every county of the State. It of course would be a rare tree throughout the rich black loam soils of the central Indiana counties. The black oak is confined to the poorer soils of the State, such as clay and gravelly ridges, sand dunes, sand ridges, and the hills of southern Indiana that are not covered with beech or white oak. It is a frequent to a common tree in the southwestern part of the State in the bottom lands where it is associated with Schneck's, shingle, and post oaks. In the northern part of the State it is generally associated with the white oak and if the soil is very poor it will form almost pure stands. On the poor ridges of southern Indiana it is generally associated with the white, and scarlet oaks, and invades habitats still poorer which are occupied by post, black jack, or chestnut oaks. Wherever the black oak is found it is generally more than a frequent tree and is usually a common tree or forms the principal stand. While the black is not so uniformly distributed over the State as the white oak, yet in point of numbers it nearly equals it, or may even exceed it.

In Floyd and Harrison Counties are certain small areas which were known to the early settlers as the "barrens." These areas were treeless. They were covered with a growth of some sort of oak which the natives call "scrub" oak, hazel, and wild plum. The height of the growth in any part would "not hide a man on horse back." These areas are now all under cultivation, and are no longer distinguished from the forested areas. However, many parts of the barrens are now covered with forests, but these forests are a complete stand of black oak. Last year one of these areas was cut off, and the age of the trees were ascertained to be about 65 years old. The barrens of southern Indiana and adjacent States offer a good problem for ecologists.

=Remarks.=--Wood similar to that of red oak, but often much inferior. The uses of the best grades of black oak are practically the same as red oak.

Where the black and scarlet oaks are associated, the scarlet oak is rarely separated from it. The two species superficially much resemble each other. The black oak is always easily distinguished by cutting into the inner bark which is yellow, while that of scarlet oak is gray or reddish. The inner bark imparts a yellow color to spittle, and the scarlet does not. When mature fruiting branches are at hand they may be separated by the appearance of the acorns. The scales of the cups of the black oak are dull, and loosely imbricated near the top while those of the scarlet oak are rather glossy and closely imbricated. The scales of the scarlet oak, however, become somewhat loose after the acorn has matured, and fallen for some time.

This species is sometimes called yellow oak. Since the chinquapin oak is also often called yellow oak, it is best to always call this species black oak.

=15.= =Quercus coccínea= Muenchhausen. Scarlet Oak. Plate 54. Medium sized trees with bark resembling the black oak, inner bark gray or reddish; twigs reddish by autumn; winter buds reddish-brown and pubescent; leaves on petioles 2.5-6 cm. long, broadly oval to obovate, blades 7-15 cm. long, truncate or wedge-shape at the base, the blade divided into 5-7 lobes by deep and wide sinuses which cut the blade more than half the distance to the midrib, sinuses rounded at the base, the lobes variable in size and shape, usually the lowest are the shortest and smallest, the middle lobes the largest and longest, the lobes widest either at the base or the apex, the terminal part toothed or lobed, the terminal lobe generally 3-lobed or 3-toothed, both surfaces of the leaves at first pubescent, soon smooth and a dark glossy green above, and paler and smooth beneath except tufts of hairs in the axils of the principal veins; acorns sessile or nearly so, solitary or in pairs; nuts ovoid to oblong, 1.5-2 cm. long, enclosed for about half their length in the thick cup-shape cup; scales triangular but blunt, closely appressed, pubescent on the back except the center which is generally elevated and smooth and shiny, giving the cup a glossy appearance which easily separates it from its nearest ally the black oak whose cup is a dull, ash or reddish gray color; kernel white within, and less bitter than the black oak.

=Distribution.=--Maine, southern Ontario to southern Nebraska, south to North Carolina, Alabama and Arkansas. It has been reported for the northwest counties and the southern part of Indiana, but we have no records for the east-central portion of the State. Clark reports it as common about Winona Lake, but does not report _Quercus velutina_ which is a common tree of the vicinity, and it is believed that Clark has confused the two species. In the northern part of the State its habitat is that of sand and gravel ridges associated with black oak. In the hill part of southern Indiana it is intimately associated with the black oak on the poorer ridges. We have no authentic records for the southwestern counties. The author has Schneck's specimens on which the record for Gibson and Posey County was based. I determined the specimens as belonging to the Spanish oak, and William Trelease verified the determination. I have no doubt that scarlet oak occurred on the sand ridges of that area.

QUERCUS COCCINEA Muenchhausen. Scarlet Oak. (× 1/2.)]

In the northern part of the State it is a rare or infrequent tree, while in favorable habitats in the hill country of the southern part of the State it is a frequent to a common tree.

=Remarks.=--Wood similar but much inferior to red oak. The cut in this State is marketed as black oak, from which it is rarely separated.

=16.= =Quercus falcàta= Michaux. Spanish Oak. Plate 55. Large trees; bark thick, rather deeply fissured, furrows usually narrow, ridges generally broad and broken into short lengths, the outer bark is reddish, except sometimes it becomes grayish by weathering; twigs densely pubescent at first, remaining more or less pubescent during the first year, or becoming smooth or nearly so and a reddish brown by autumn; leaves on petioles 0.5-6 cm. long, ordinarily about 2-3 cm. long, blades very variable in outline, ovate, ovate-oblong or obovate, usually somewhat curved, wedge-shaped, rounded or truncate at the base, shallow or deeply lobed, generally about 2/3 of the distance to the midrib; lobes 3-11, commonly 5-9, the number, size and shape of the lobes exceedingly variable, the longest lateral lobes are generally near the middle of the leaf, sometimes the lowest pair, sometimes the upper pair are the longest, terminal lobe triangular or oblong, generally widest at the base, although frequently widest at the apex, lateral lobes widest at the base and gradually becoming narrower, towards the apex, rarely somewhat wider at the apex, generally somewhat curved, lobes generally sharp-pointed, sometimes wide-angled or rounded at the apex, margins of lobes entire, wavy, toothed or lobed, sinuses wide and rounded at the base; leaves densely pubescent on both surfaces at first, gradually becoming smooth and dark green above by autumn, the under surface remaining covered with a tomentum which is grayish or yellowish; acorns sessile or nearly so, solitary or in pairs; nuts broadly ovoid, generally 10-12 mm. long, broadly rounded at the base, rounded at the apex, enclosed about one-half their length by the cup; cups strongly convex at the base; scales blunt, grayish and pubescent on their backs, their margins reddish and generally smooth.

=Distribution.=--New Jersey and Missouri, south to Florida and west to Texas. The known distribution in Indiana would be that part of the State south of a line drawn from Vincennes to North Madison. It is local except in the southwestern counties. In our area it is found on both high and low ground. In Jefferson and Clark Counties it is found only in the flats where it is associated with beech, sweet gum, pin oak, red maple and black gum. A colony was found in Washington County on high ground, about eight miles southwest of Salem associated with black and post oak. In Harrison County about two miles southeast of Corydon it was found on the crest of a ridge with white and black oak. In Daviess County about four miles east of Washington it is associated with black and post oak. In Knox, Gibson, Pike and Warrick Counties it is local on sand ridges with black oak. It occurs in the greatest abundance in the river bottoms of Gibson, Posey and Spencer Counties, where it is generally associated with pin, Schneck's, shingle, swamp white, black and post oaks, and sweet gum. In the last named counties it is fairly well distributed, and is a frequent to a common tree. Brown's[43] report for Fountain County should not be recognized without a verifying specimen, since his list was compiled from a list of common names of the trees which he obtained.

QUERCUS FALCATA Michaux. Spanish Oak. (× 1/2.)]

=Remarks.=--Wood and uses similar to that of red oak. In Indiana it is all sold as red oak. In all parts of its range in Indiana it is known as red or black oak. However, the best accepted common name of this species throughout its range is Spanish oak, and since no other species is known by this name, it should be used for this species.

The bark of this species varies considerably in color and tightness. The leaves are exceedingly variable in form. The leaves on the same tree will vary from 3-lobed to 11-lobed. Usually the lobing is deepest in the leaves nearest the top of the tree. Leaves of small trees, coppice shoots, and of the lower branches of some trees are often all or for the greater part 3-lobed. The color of the pubescence of the lower surface of the leaves varies from a gray to a yellow-gray. The variations have lead authors to divide this polymorphic species into several species and varieties. The author has included all the forms that occur in Indiana under one name.

This species is variously known as _Quercus digitata_, _Quercus triloba_, _Quercus pagodaefolia_, and by the most recent authors as _Quercus pagoda_ and _Quercus rubra_ and its varieties. Specimens in the author's collection from Jefferson County were reported by Sargent[44] as _Quercus rubra_ var. _triloba_.

=17.= =Quercus marilándica= Muenchhausen. Black Jack Oak. Plate 56. Mature trees generally 10-30 cm. in diameter; bark resembles that of a gnarled black oak; twigs generally scurvy-pubescent the first year; leaves on petioles from nearly sessile to 2.5 cm. long, usually less than a cm. long, blades 7-15 cm. long, broadly obovate, often almost as wide as long, narrowly rounded at the base, with three primary lobes at the apex, sometimes with two small lateral lobes, the apex is sometimes almost rounded and the position where the lobes usually occur is indicated by three primary veins which end in a bristle, the apex of the leaf is generally about equally divided into three lobes by two very shallow rounded sinuses, the lobes are rounded or merely acute; sometimes the terminal lobes develop a secondary lobe, leaves very pubescent both above and beneath when they first appear, becoming smooth and glossy above at maturity, and remaining more or less pubescent beneath; acorns sessile or nearly so, single or in pairs; nuts ovoid or oblong, 1-1.5 cm. long, broadly rounded at the base, rounded or somewhat conic at the apex, enclosed for about half their length in the cup-shaped cup; scales blunt, not closely appressed, pubescent on back, light reddish-brown; kernel bitter.

QUERCUS MARILANDICA Muenchhausen. Black Jack Oak. (× 1/2.)]

=Distribution.=--New York to Nebraska, south to Florida and west to Texas. In Indiana it is known to the author from Sullivan, Greene and Clark Counties and southwestward. It has been reported from Jefferson County by Barnes which is no doubt correct. Doubtful records are those by Brown for Fountain County, Miami County by Gorby, and Phinney's report for the area of Delaware, Jay, Randolph and Wayne Counties. It has been reported for the vicinity of Chicago by Higley and Raddin. It may be local on sterile, sandy ridges of the northern part of the State, but very local if it does occur. It is generally found in very poor soil on the crest of ridges associated with black and post oak. However, it has been found in Greene, Sullivan and Knox counties on sand ridges and at the base of sand ridges associated with black and post oak. The species has a very limited mass distribution and is only occasionally found and in colonies of a few trees each.

=Remarks.=--Trees too small and scarce to be of any economic importance.

ULMÀCEAE. The Elm Family.

Trees or shrubs with simple, alternate, 2-ranked, petioled leaves; sepals 3-9, petals none, stamens as many as the sepals and opposite them, stigmas 2.

Branchlets with solid pith; leaves with primary veins
parallel; flowers borne on the twigs of the preceding
season 1 Ulmus.

Branchlets with chambered pith at the nodes; leaves
3-veined at the base; flowers borne on the twigs of
the season 2 Celtis.

1. ÚLMUS. The Elms.

Trees with furrowed bark; leaves short petioled, with lateral veins prominent and parallel, oblique or unequally heart-shaped at the base, taper-pointed at the apex, mostly double-serrate; flowers of Indiana species expanding before the leaves in March or April; fruit a samara surrounded with a wide membranous margin, maturing in the spring.

Inner bark mucilaginous; leaves very rough above; flowers
nearly sessile; fruit not ciliate 1 U. fulva.

Inner bark not mucilaginous; leaves smooth or somewhat rough
above; flowers on slender pedicils; fruit ciliate.

Branches without corky wings; sides of samara glabrous 2 U. americana.

Branches (at least some of them) with corky wings; at least
one side of the samara pubescent.

Buds ovate, not twice as long as wide, obtuse, or
short-pointed, dark brown; scales pubescent and
ciliate; leaves usually not twice as long as wide,
base of petiole glabrous beneath; calyx lobes 7-9 3 U. Thomasi.

Buds small, narrow, twice as long as wide, very
sharp-pointed, light brown; scales glabrous or
merely puberulent; leaves usually twice as long
as wide, base of petiole pubescent all around 4 U. alata.

=1. Ulmus fúlva= Michaux. Slippery Elm. Red Elm. Plate 57. Fairly large trees with deeply fissured reddish-brown bark without white streaks between the layers of the ridges, twigs very pubescent and green at first, becoming gray or reddish-brown at the end of the season and remaining more or less pubescent for a year or more; buds ovate, a very dark reddish brown, the scales more or less pubescent; leaves ovate, oval or slightly obovate, average blades 8-15 cm. long, hairy on both surfaces at first, remaining more or less pubescent beneath until maturity, and becoming very rough above with a few scattered hairs remaining, fragrant when dried, fragrance remaining for years; fruit ripening the last of April or the first of May before or with the unfolding of the leaves; samara orbicular or obovate, usually longer than wide, average size 13-17 mm. long and 9-12 mm. wide, the margin as wide or wider than the seed, margin glabrous, seed densely pubescent on both sides; wood hard, strong, light when well seasoned and not warping as badly as white elm.

=Distribution.=--Quebec south to Florida, west to Texas, Nebraska and North Dakota. Found in all parts of Indiana. In the prairies or in the "flats" it may be absent in one or more contiguous counties and may be entirely absent on the crests and upper slopes of ridges. It prefers a moist well drained soil, and where it is found it is usually a frequent to a common tree, although rarely is it found as a very common tree. It is usually associated with sugar maple, beech, white ash, linn, tulip, white oak, etc.

=Remarks.=--This tree usually is from 3-6 dm. in diameter and tall for its diameter. However, larger trees occur. In the Ind. Geol. Rept. 6:70:1875 mention is made of a tree in Jackson County that was "18 feet in circumference." The uses of the wood are similar to that of white elm. The inner bark collected in spring is much used in medicine under the name of slippery elm.

ULMUS FULVA Michaux. Red or Slippery Elm. (× 1/2.)]

=2. Ulmus americàna= Linnæus. White Elm. Plate 58. Large trees; bark deeply fissured, gray, the ridges showing white streaks between the layers; twigs more or less hairy at first and usually becoming glabrous by the end of the season; buds ovate, acute and glabrous; leaves ovate, oval or obovate, average blades 8-12 cm. long, hairy on both sides on expanding, becoming at maturity glabrous above and smooth or rough, sometimes very rough on vigorous young shoots, remaining pubescent beneath, rarely glabrous; fruit ripening before or as the leaves unfold, generally oval in shape, about 1 cm. long, both surfaces glabrous, margins about as wide as the seed and fringed with hairs; wood hard, tough, flexible, generally hard to split, warps badly in seasoning.

=Distribution.=--Quebec to Florida, west to Texas and Nebraska. Found throughout Indiana, and doubtless in every county. It is frequent to common or very common on the flood plains of streams, in wet woods and in low ground generally.

=Remarks.=--This species is also called water elm, swamp elm, gray elm, bitter elm, sour elm and in southwestern counties it is often called red elm. In Perry County it is often called hub elm. It is generally known as "elm" and when this term is used, it refers to this species. The wood has a very wide range of uses. The greatest amount has been used for hoops, staves and heading. Large quantities have been used in the manufacture of agricultural implements, hubs, furniture, basket handles, etc. White elm is usually considered very difficult to split, but I was informed by a pioneer timber cutter that the heart wood of the veterans of the forest splits as well as oak, and that he worked many a tree up into staves. He told me that he made into staves a tree in Paulding County, Ohio, that was eight feet in diameter at the stump. There is little attempt being made by woodlot owners to propagate this tree. However, the natural propagation of the species is probably greater than any other species because it produces seed at an early age, and culls of the forest are not cut because they are not good for fuel which leaves them to produce seed. Then the seed are light, and are scattered to great distances by the wind and water. It is propagated very easily from seedlings.

The tree when grown in the open has a tendency to be bushy and unless it is given some pruning will have a very short clear trunk. It has always been regarded as one of the best species for shade tree planting. For beauty of form it is not excelled by any tree for shade or ornamental planting. However, it has several insect enemies that require spraying to keep them under control.

ULMUS AMERICANA Linnæus. White Elm. (× 1/2.)]

=3. Ulmus Thomási= Sargent. Hickory Elm. Rock Elm. Plate 59. Large trees; bark deeply fissured and grayish like the bark of the white elm; twigs light brown, generally densely hairy and remaining more or less pubescent until the end of the season or later, the twigs of some specimens are glabrous or only slightly hairy at first and soon become glabrous and somewhat glaucous, after the first year some of the branchlets begin to develop 1-4 corky ridges from a few millimeters to 5 or 6 mm. in thickness, the ridges are wide and rounded at the top, dark gray, brown and discontinuous, rarely a corky ridge will appear on a branchlet the first year; leaves oval or obovate, average blades 8-15 cm. long, at maturity glabrous and smooth or rough to very rough above, permanently pubescent beneath especially on the veins; fruit ripens late in May or early in June when the leaves are from 1/2 to 2/3 grown; samara oval, usually 1.5-2 cm. long, oblique at the base, with a beak 2-5 mm. long at the apex, both faces pubescent, wing about as wide as the seed; wood hard heavy, strong, flexible, uses the same as white elm.

=Distribution.=--Southern Quebec and Ontario south to northern New Jersey and west to Minnesota and Missouri. The distribution in Indiana has not been studied. The frequency of its occurrence is not known, and all of the known stations are given. The published records are as follows: Dearborn (Collins); Franklin (Meyncke); Hamilton (Wilson); Jefferson (Barnes) and (Deam); Noble (VanGorder); Parke (Hobbs); St. Joseph (Nieuwland); Steuben (Bradner); Wayne (Petry and Markle); Wells (Deam). Additional records are Hendricks, Noble, Ripley, Vermillion and Wayne by Deam. The published record for Posey County by Deam and Schneck should be referred to _Ulmus alata_. It prefers a well drained soil and is most frequently found near the base of the slope or on the top of flood plain banks of streams, in ravines, or in a habitat like a beech-sugar maple woods. It is reported to have been frequent in Franklin, Noble and Wells Counties.

Its appearance and habit of growth is so much like the white elm that it is not commonly distinguished from it, which accounts for the lack of definite knowledge of its range in our area.

ULMUS THOMASI Sargent. Hickory or Rock Elm. (× 1/2.)]

ULMUS ALATA Michaux. Winged Elm. (× 1/2.)]

=4. Ulmus alàta= Michaux. Winged Elm. Plate 60. Small to medium sized trees; bark rather closely fissured, grayish or reddish-brown, in appearance like white elm; twigs hairy at first, generally remaining more or less pubescent throughout the season, rarely becoming entirely glabrous before the end of the season, a light brown gradually becoming a gray-brown; branchlets usually begin to develop two thin narrow corky ridges, becoming by the end of the second year 4-7 mm. thick, the year's growth of corky layer a light brown, the older layers a darker brown, the two main corky ridges are on opposite sides of the twigs, and between these there are generally additional corky excrescences, especially on the older branches; leaves oblong-lanceolate or oval, some somewhat falcate, average blades 4-8 cm. long, pubescent on both sides on unfolding, becoming at maturity glabrous or nearly so above, some are rough above at maturity, remaining pubescent until maturity beneath; petioles short, generally 2-3 mm. long, rarely 5 mm. or longer; fruit ripening before or with the unfolding of the leaves; samara 6-10 mm. long, pubescent on both faces.

=Distribution.=--Virginia west through southern Indiana to southern Missouri, south to the Gulf and west to Texas. In Indiana it is confined to the southwestern part of the State. Gorby's report for Miami should be ignored. It has been reported as far north as Vigo and Monroe Counties by Blatchley, and as far east as Clark County by Baird and Taylor. The author has collected it in Crawford, Dubois, Martin, Orange, Perry, Posey, Spencer and Warrick Counties.

The tree has two rather distinct habitats. In the hill counties it is found on the sides of cliffs, steep slopes or on the top of the ridges with such species as the black, chestnut and scarlet oaks and chestnut. In this habitat it is usually a small scrubby tree with an excessive number of side branches. Such specimens usually have the corky ridges well developed on all of the branches and the tree presents a weird appearance. The second habitat is in the hard clay flats of the southwestern counties. In Warrick County along Big Pigeon Creek west of Boonville I measured a specimen 21 dm. in circumference and I estimated the clear bole at 8 m. It was associated with sweet gum, black gum, white elm, red birch, red oak, etc. It is found throughout this county both in the "flats" and on the sandy ridges. In Posey County it is a frequent tree in the low woods about 10 miles southwest of Mt. Vernon. In these woods it acquires a diameter of 3-6 dm. and is associated with post oak, Spanish oak, sweet gum, shingle oak, etc. It is to be noted that specimens that grow in these conditions and those that acquire a large size do not develop such conspicuous corky branches. A large tree over 6 dm. in diameter was noted in the eastern part of Gibson County growing in low sandy soil which was destitute of corky branches so far as could be seen from the ground. All of the branches examined were free from corky ridges, and only a few corky excrescences were present. The specimen could easily be identified by the leaves. Another large tree 12 dm. in circumference in a black oak woods 4 miles south of Marengo in Crawford County was also free from corky ridges.

This is an interesting tree and requires further study to establish its range in Indiana and to learn its habits. In Jasper, Indiana, it is a frequent shade tree. No doubt the trees were obtained from a nearby woods along the Patoka River where this species is known to occur.

=2. CÉLTIS.= The Hackberries.

Trees with pith of branchlets chambered; flowers in Indiana species appear before the leaves, the leaves generally with 3 primary veins at the base; staminate flowers usually in clusters, the pistillate solitary or few together in the axils of the leaves, and near the end of the twigs; fruit a globose drupe, sometimes elongated, pulp thin and sweet, frequently remaining on the tree until late winter, relished by birds; stone bony, wrinkled.

Some of the American species of hackberry are very variable. The habitat of the species varies from deep swamps to arid rocky slopes. In fact, a single species as now understood may have a variable habitat. The following variations may be noted on the same tree or on different trees of the same species. The twigs may be glabrous, or pubescent; the leaves may vary in size, shape and texture, leaf margin, and in the roughness or smoothness of the surfaces; the petioles may be smooth or hairy; the pedicels may be glabrous or pubescent, shorter or longer than the petioles; the fruit also varies in shape. Leaves have been seen on the same tree which were smooth above, while others were quite rough above, the difference being due to the exposure to light.

The original descriptions of the species are too short to sufficiently characterize the species, which adds to the confusion. However, C. S. Sargent[45] has recently revised the species and varieties of our area.

Prof. Sargent has examined and named all of my material for me. Mr. B. F. Bush, who has extensively studied the hackberries in the field, also has examined my specimens.

The writer has paid special attention to the hackberries of the State for the past few years and is still in doubt as to the status of the species that occur in the State. Since I am not following the determinations made by Sargent and Bush, and am following the nomenclature of the first edition, I regard the present treatment as tentative only.

Margins of all the leaves sharply serrate all around
except at base; nutlets 6-8 mm. long 1 C. occidentalis.

Margins of leaves of fruiting branches generally entire,
or some with a few teeth on one side or with a few
teeth on both sides; margins of the leaves of
vegetative branches and shoots similar to those of
fruiting branches or with the margins serrate nearly
all around; nutlets 5-6 mm. long.

Leaves of a rather broad ovate type; mature fruit a
dark cherry-red; usually shrubs, sometimes very
small trees, of a dry habitat 2 C. pumila.

Leaves of an ovate-lanceolate type; mature fruit a
light cherry-red; medium-sized trees of a wet
habitat 3 C.
mississippiensis.

CELTIS OCCIDENTALIS Linnæus. Hackberry. (x 1/2.)]

=1. Celtis occidentàlis= Linnæus. Hackberry. Plate 61. Medium to large-sized trees; bark of old trees irregularly furrowed, sometimes some of the surface warty and rough; twigs smooth or pubescent, the fruiting ones generally smooth; leaves of an ovate type on petioles 0.5-2 cm. long, the blades of fruiting twigs 5-15 cm. long, those of vegetative twigs sometimes larger, oblique or slightly cordate at base, gradually tapering to a point at apex, or long acuminate at the apex, often becoming thick at maturity, especially those exposed to full sunlight, generally smooth above at maturity, especially those of fruiting twigs, or sometimes rough, especially those of vegetative branchlets or those growing in the shade, the under surface more or less pubescent along the veins at maturity; fruit matures in late autumn, very dark red, sometimes appearing almost black, globose or somewhat oblong, generally about 9-10 mm. in diameter, borne on pedicels which are longer or up to twice as long as the petioles; the pedicels which are always ascending are straight or somewhat curved upwards; nutlets globose, a little longer than wide.

=Distribution.=--Valley of the St. Lawrence River, southern Ontario, to North Dakota, and south to the Gulf States and west to Texas. More or less frequent along streams throughout the State, except in the hilly counties of the southern part of the State. It is always found in moist soil, except in the hilly counties where it may be found on wooded slopes or on high rocky bluffs bordering streams. In all of our area the species is practically confined to drainage basins, and is generally close to streams.

=Remarks.=--The wood is yellowish-white and before seasoning very much resembles ash for which it was generally sold. It has good bending qualities and is now much sought after for hoops. It was formerly often known as hoop ash. The supply is now becoming scarce, but when bought sells for the same price as good white elm.

Some writers include under the name _Celtis occidentalis_ only those forms which are small trees and have ovate, short-pointed leaves. This type of tree has not been found in Indiana. The form with long acuminate pointed leave which is the common form in our area, is regarded as a variety of _Celtis occidentalis_. Trees having the upper surface of the leaves very rough are called _Celtis crassifolia_ Lamarck, or are merely regarded as a variety of _Celtis occidentalis_. This form is found throughout our area.

The hackberry is sometimes used as a shade tree. It can scarcely be recommended because its leaves and twigs are often affected by galls which detract from its appearance.

CELTIS PUMILA var. DEAMII Sargent. Dwarf Hackberry. (× 1/2.)]

=2. Celtis pùmila= (Muhlenberg) Pursh. Dwarf Hackberry. Plate 62. Bark thin, smooth and gray on shrub-like forms, warty or deeply fissured on the larger forms; ridges flat and broken, dark gray-brown; twigs at first hairy, becoming smooth or nearly so by autumn; leaves of an ovate type, broadly-ovate, oblong-ovate to narrow ovate, on petioles 0.5-1.5 cm. long, blades of fruiting branchlets 3-10 cm. long, those of sterile twigs sometimes larger, oblique, rounded or somewhat cordate at the base, taper-pointed, sometimes acuminate at the apex, margins entire or with a few teeth usually about or above the middle, becoming thick and smooth above at maturity, sometimes rough, especially on vigorous shoots, generally somewhat pubescent along the veins beneath; fruit matures late in the autumn, usually an orange or light cherry color late in summer, becoming a very dark cherry color late in the autumn, globose to ellipsoidal, on pedicels about as long as the petioles; sometimes the pedicels are shorter but usually about one-half longer; pedicels generally ascending, rarely recurved, when recurved the pedicels are short.

=Distribution.=--Pennsylvania to northern Illinois, south to Florida and west to Arkansas. Local in Indiana. It has been collected by the writer in Lake County near the mouth of the Grand Calumet River where it was collected by E. J. Hill who has given us the most detailed account of this species.[46] Also collected on a high, gravelly hill on the east side of Hog-back Lake, Steuben County; on a rocky wooded slope in Hamar's Hollow southeast of Mitchell in Lawrence County; on a "knob" in Floyd County; on a rocky wooded slope near Big Spring in Washington County; frequent on a rocky wooded slope near the Ohio River east of Elizabeth in Harrison County; on the bank of Blue River near Milltown in Crawford County; and in Perry County along the bluffs of the Ohio River about six miles east of Cannelton, and also on the crest of a ridge about six miles southwest of Derby. It has also been reported by Nieuwland for Clark in Marshall County.

=Remarks.=--This species is usually a small shrub, and usually bears fruit when only 1.5-2 meters (5 or 6 feet) tall. Only a few trees have been seen that were 40 cm. (4 inches) in diameter. The small size at which this species fruits, easily distinguishes it from other species in our area. Its habitat also serves to distinguish it. Along Lake Michigan it grows on the dry sand dunes, and in southern Indiana it grows on dry rocky slopes.

Sargent who has examined all of my specimens credits Indiana with the typical species, and separates from it a form which he calls _Celtis pumila_ variety _Deamii_[47]. This variety is based upon my No. 18,727, and the type specimen has been photographed to illustrate this species. The writer is not able to separate the two forms in our area, and believes that all belong either to _Celtis pumila_ or to the new variety.